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The ocean is teeming with a myriad of mm-sized invertebrate planktonic larvae, which thrive in a viscous fluid environment. Many of them rely on ciliary beating to generate fluid flows for locomotion and feeding. Their larval forms, local…

We derive from first principles a three-dimensional theory of self-propelled particle swarming in a viscous fluid environment. Our model predicts emergent collective behavior that depends critically on fluid opacity, mechanism of…

Soft Condensed Matter · Physics 2026-05-12 Yao-Li Chuang , M. R. D'Orsogna , T. Chou

The Physarum network expands or retracts in response to environmental stimuli, demonstrating an intelligent adaptive capability to locate optimal paths for nutrient transport. The underlying physical mechanism governing this intelligence…

Biological Physics · Physics 2026-02-03 Bingyang Han , Luolan Chen , Tieyan Si

Biological membranes are host to proteins and molecules which may form domain-like structures resulting in spatially-varying material properties. Vesicles with such heterogeneous membranes can exhibit intricate shapes at equilibrium and…

Soft Condensed Matter · Physics 2023-06-22 Prerna Gera , David Salac , Saverio E. Spagnolie

Tissue remodelling in diverse developmental contexts require cell shape changes that have been associated with pulsation and flow of the actomyosin cytoskeleton. Here we describe the dynamics of the actomyosin cytoskeleton as a confined…

Biological Physics · Physics 2016-05-25 Deb Sankar Banerjee , Akankshi Munjal , Thomas Lecuit , Madan Rao

A big challenge in current biology is to understand the exact self-organization mechanism underlying complex multi-physics coupling developmental processes. With multiscale computations of from subcellular gene expressions to cell…

Cell Behavior · Quantitative Biology 2021-05-18 Xiaoliang Wang , Dongyun Bai

Living organisms are molecular systems with self-sustained dynamics via energy conversion through molecular cooperation, resulting in highly complex macroscopic behaviors. Construction of such autonomous macroscopic dynamics at a molecular…

We investigate the hovering dynamics of rigid bodies with up-down asymmetry placed in oscillating background flows. Recent experiments on inanimate pyramid-shaped objects in oscillating flows with zero mean component demonstrate that the…

Fluid Dynamics · Physics 2016-09-21 Yangyang Huang , Monika Nitsche , Eva Kanso

A great number of biological organisms live in aqueous environments. Major evolutionary transitions, including the emergence of life itself, likely occurred in such environments. While the chemical aspects of the role of water in biology…

Populations and Evolution · Quantitative Biology 2020-07-01 Madison S. Krieger , Sam Sinai , Martin A. Nowak

We review the dynamical behavior of giant fluid vesicles in various types of external hydrodynamic flow. The interplay between stresses arising from membrane elasticity, hydrodynamic flows, and the ever present thermal fluctuations leads to…

Soft Condensed Matter · Physics 2014-05-12 David Abreu , Michael Levant , Victor Steinberg , Udo Seifert

Active crystals are highly ordered structures that emerge from the self-organization of motile objects, and have been widely studied in synthetic and bacterial active matter. Whether collective crystallization phenomena can occur in groups…

We study the spatial distribution of gyrotactic microorganisms transported by a three-dimensional turbulent flow generated by direct numerical simulations. We find that gyrotaxis combines with turbulent fluctuations to produce small scales…

Chaotic Dynamics · Physics 2012-06-13 Filippo De Lillo , Guido Boffetta , Massimo Cencini

When interacting motile units self-organize into flocks, they realize one of the most robust ordered state found in nature. However, after twenty five years of intense research, the very mechanism controlling the ordering dynamics of both…

Soft Condensed Matter · Physics 2021-10-04 Amélie Chardac , Ludwig A. Hoffmann , Yoann Poupart , Luca Giomi , Denis Bartolo

We use numerical simulations to systematically investigate the vesicle dynamics in two-dimensional (2D) Taylor-Green vortex flow in the absence of inertial forces. Vesicles are highly deformable membranes encapsulating an incompressible…

Biological Physics · Physics 2023-05-10 Gökberk Kabacaoğlu , Enkeleida Lushi

This study investigates the three-dimensional organisation and evolution of stall cells in the separated flow region over an airfoil. Using a hybrid RANS/LES approach based on the DDES-SST turbulence model, we characterise the formation and…

Fluid Dynamics · Physics 2026-02-11 Rishabh Mishra , Emmanuel Guilmineau , Ingrid Neunaber , Caroline Braud

Active fluids exhibit spontaneous flows with complex spatiotemporal structure, which have been observed in bacterial suspensions, sperm cells, cytoskeletal suspensions, self-propelled colloids, and cell tissues. Despite occurring in the…

Soft Condensed Matter · Physics 2022-10-05 Ricard Alert , Jaume Casademunt , Jean-François Joanny

The complex spatiotemporal flow patterns in living tissues, driven by active forces, have many of the characteristics associated with inertial turbulence even though the Reynolds number is extremely low. Analyses of experimental data from…

Soft Condensed Matter · Physics 2022-11-29 Xin Li , Sumit Sinha , T. R. Kirkpatrick , D. Thirumalai

Geometric confinement plays an important role in the dynamics of natural and synthetic microswimmers from bacterial cells to self-propelled particles in high-throughput microfluidic devices. However, little is known about the effects of…

Fluid Dynamics · Physics 2014-11-13 Alan Cheng Hou Tsang , Eva Kanso

Cytoplasmic streaming, the persistent flow of fluid inside a cell, induces intracellular transport, which plays a key role in fundamental biological processes. In meiosis II mouse oocytes (developing egg cells) awaiting fertilisation, the…

Biological Physics · Physics 2024-10-18 Weida Liao , Eric Lauga

Here we show that coupling to curvature has profound effects on collective motion in active systems, leading to patterns not observed in flat space. Biological examples of such active motion in curved environments are numerous: curvature…

Soft Condensed Matter · Physics 2015-03-05 Rastko Sknepnek , Silke Henkes