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Sperm traverse their microenvironment through viscous fluid by propagating flagellar waves; the waveform emerges as a consequence of elastic structure, internal active moments, and low Reynolds number fluid dynamics. Engineered…

Fluid Dynamics · Physics 2015-02-19 Thomas Montenegro-Johnson , Hermes Gadelha , David J. Smith

Dense suspensions of swimming bacteria are known to exhibit collective behaviour arising from the interplay of steric and hydrodynamic interactions. Unconfined suspensions exhibit transient, recurring vortices and jets, whereas those…

Biological Physics · Physics 2016-07-04 Hugo Wioland , Enkeleida Lushi , Raymond E. Goldstein

The motion of several self-propelled boats in a narrow channel displays spontaneous pattern formation and kinetic phase transitions. In contrast with previous studies on self-propelled particles, this model does not require stochastic…

Statistical Mechanics · Physics 2012-06-13 Eric Heisler , Nobuhiko J. Suematsu , Akinori Awazu , Hiraku Nishimori

We study the emerging collective states in a simple mechanical model of a dense group of self-propelled polar disks with off-centered rotation, confined within a circular arena. Each disk presents self-alignment towards the sum of contact…

Soft Condensed Matter · Physics 2026-04-15 Weizhen Tang , Yating Zheng , Amir Shee , Guozheng Lin , Zhangang Han , Pawel Romanczuk , Cristián Huepe

We present the hydrodynamic theory of coherent collective motion ("flocking") at a solid-liquid interface, and many of its predictions for experiment. We find that such systems are stable, and have long-range orientational order, over a…

Soft Condensed Matter · Physics 2022-01-05 Niladri Sarkar , Abhik Basu , John Toner

Stall cells are spanwise-periodic flow structures that spontaneously form on airfoils operating near stall, fundamentally altering the aerodynamic loading distribution. Despite decades of experimental observations, a complete theoretical…

Fluid Dynamics · Physics 2026-02-11 Rishabh Mishra , Emmanuel Guilmineau , Ingrid Neunaber , Caroline Braud

Meso-scale turbulence was originally observed experimentally in various suspensions of swimming bacteria, as well as in the collective motion of active colloids. The corresponding large-scale dynamical patterns were reproduced in a simple…

Soft Condensed Matter · Physics 2021-03-30 Vasco M. Worlitzer , Gil Ariel , Avraham Be'er , Holger Stark , Markus Bär , Sebastian Heidenreich

This work focuses on the formation of mating aggregates in zooplankton. In particular, sexual encounters are behaviourally supported by males actively swimming in search for females, and approaching them for mating once they are found.…

Populations and Evolution · Quantitative Biology 2022-05-19 Ron Shnapp , François-Gaël Michalec , Markus Holzner

We introduce a stochastic agent-based model for the flocking dynamics of self-propelled particles that exhibit velocity-alignment interactions with neighbours within their field of view. The stochasticity in the dynamics of the model arises…

Statistical Mechanics · Physics 2019-07-24 Trilochan Bagarti , Shakti N. Menon

We study clustering and percolation phenomena in the Vicsek model, taken here in its capacity of prototypical model for dry aligning active matter. Our results show that the order-disorder transition is not related in any way to a…

Soft Condensed Matter · Physics 2019-09-04 Nikos Kyriakopoulos , Hugues Chaté , Francesco Ginelli

A spacially extended model of the collective behavior of a large number of locally acting organisms is proposed in which organisms move probabilistically between local cells in space, but with weights dependent on local morphogenetic…

adap-org · Physics 2008-06-25 Mark M. Millonas

Self-propulsion of cellular microswimmers generates flow signatures, commonly classified as pusher- and puller-type, which characterize hydrodynamic interactions with other cells or boundaries. Using experimentally measured beat patterns,…

Cell Behavior · Quantitative Biology 2016-01-20 Gary S. Klindt , Benjamin M. Friedrich

Understanding the organization of collective motion in biological systems is an ongoing challenge. In this Paper we consider a minimal model of self-propelled particles with variable speed. Inspired by experimental data from schooling fish,…

Biological Physics · Physics 2015-06-04 Shradha Mishra , Kolbjørn Tunstrøm , Iain D. Couzin , Cristián Huepe

A hallmark in natural systems, self-organization often stems from very simple interaction rules between individual agents. While single-species self-propelled particle (SPP) systems are well understood, the behavior of binary mixtures with…

Soft Condensed Matter · Physics 2026-03-05 Eloise Lardet , Letian Chen , Thibault Bertrand

We introduce a model for self-organized dynamics which, we argue, addresses several drawbacks of the celebrated Cucker-Smale (C-S) model. The proposed model does not only take into account the distance between agents, but instead, the…

Analysis of PDEs · Mathematics 2015-05-27 Sebastien Motsch , Eitan Tadmor

Bacteria exist in a free-swimming state or in a sessile biofilm state. The transition from free-swimming to sessile mode is characterized by changes in gene expression which alter, among others,the mechanics of flagellar motility. In this…

Fluid Dynamics · Physics 2014-05-07 Alan Cheng Hou Tsang , Eva Kanso

A type of polar self-propelled particle generates a torque that makes it naturally drawn to higher-density areas. The collective behaviour this induces in assemblies of particles constitutes a new form of phase separation in active fluids.

Soft Condensed Matter · Physics 2021-05-24 Olivier Dauchot

Active matter consists of units that generate mechanical work by consuming energy. Examples include living systems, such as assemblies of bacteria and biological tissues, biopolymers driven by molecular motors, and suspensions of synthetic…

Soft Condensed Matter · Physics 2021-02-04 Song Liu , Suraj Shankar , M. Cristina Marchetti , Yilin Wu

The aim of the present paper is to elucidate the transition from collective to random behavior exhibited by various mathematical models of bird flocking. In particular, we compare Vicsek's model [Viscek et al., Phys. Rev. Lett. 75, 1226 --…

Adaptation and Self-Organizing Systems · Physics 2015-06-17 H. Christodoulidi , K. van der Weele , Ch. G. Antonopoulos , T. Bountis

Suspensions of swimming micro-organisms provide examples of coordinated active dynamics. That has stimulated the study of a phenomenological theory combining synchronization and polar order in active matter. Here, we consider another…

Soft Condensed Matter · Physics 2019-04-08 M. Leoni