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The self-organised motion of vast numbers of creatures in a single direction is a spectacular example of emergent order. We recreate this phenomenon using actuated non-living components. We report here that millimetre-sized tapered rods,…

Soft Condensed Matter · Physics 2014-09-23 Nitin Kumar , Harsh Soni , Sriram Ramaswamy , A. K. Sood

Systems composed of strongly interacting self-propelled particles can form a spontaneously flowing polar active fluid. The study of the connection between the microscopic dynamics of a single such particle and the macroscopic dynamics of…

Soft Condensed Matter · Physics 2018-03-02 Benjamin Loewe , Anton Souslov , Paul M. Goldbart

We have recorded the swarming-like collective migration of a large number of keratocytes (tissue cells obtained from the scales of goldfish) using long-term videomicroscopy. By increasing the overall density of the migrating cells, we have…

Cell Behavior · Quantitative Biology 2009-11-13 Balint Szabo , Gergely J. Szollosi , Balazs Gonci , Zsofi Juranyi , David Selmeczi , Tamas Vicsek

The collective motion of epithelial cells is a fundamental biological process which plays a significant role in embryogenesis, wound healing and tumor metastasis. While it has been broadly investigated for over a decade both in vivo and in…

Life in complex systems, such as cities and organisms, comes to a standstill when global coordination of mass, energy, and information flows is disrupted. Global coordination is no less important in single cells, especially in large oocytes…

We study a novel phase of active polar fluids, which is characterized by the continuous creation and destruction of dense clusters due to self-sustained turbulence. This state arises due to the interplay of the self-advection of the aligned…

Soft Condensed Matter · Physics 2022-07-12 Vasco M. Worlitzer , Gil Ariel , Avraham Be'er , Holger Stark , Markus Bär , Sebastian Heidenreich

We extensively study the Toner-Tu-Swift-Hohenberg model of motile active matter by means of direct numerical simulations in a two-dimensional confined domain. By exploring the space of parameters of the model we investigate the emergence of…

Fluid Dynamics · Physics 2023-06-02 Leonardo Puggioni , Guido Boffetta , Stefano Musacchio

The complex formations exhibited by schooling fish have long been the object of fascination for biologists and physicists. However, the physical and sensory mechanisms leading to organized collective behavior remain elusive. On the physical…

Fluid Dynamics · Physics 2025-07-09 Anand U. Oza , Eva Kanso , Michael J. Shelley

The collective behaviour of confluent cell sheets is strongly influenced both by polar forces, arising through cytoskeletal propulsion and by active inter-cellular forces, which are mediated by interactions across cell-cell junctions. We…

Soft Condensed Matter · Physics 2020-05-28 Guanming Zhang , Romain Mueller , Amin Doostmohammadi , Julia M. Yeomans

We derive from first principles a three-dimensional theory of self-propelled particle swarming in a viscous fluid environment. Our model predicts emergent collective behavior that depends critically on fluid opacity, mechanism of…

Soft Condensed Matter · Physics 2026-05-12 Yao-Li Chuang , M. R. D'Orsogna , T. Chou

Active swimmers are ubiquitous in nature, found in many diverse biological systems ranging from bacteria to vertebrate fish. Of particular importance are sperm cells which are swimmers that are crucial for the survival of many species…

Biological Physics · Physics 2017-11-15 James L. Kingsley , Utkan Demirci , Erkan Tuzel

A wide range of experimental systems including gliding, swarming and swimming bacteria, in-vitro motility assays as well as shaken granular media are commonly described as self-propelled rods. Large ensembles of those entities display a…

Statistical Mechanics · Physics 2021-01-18 Markus Bär , Robert Großmann , Sebastian Heidenreich , Fernando Peruani

We construct the hydrodynamic theory of coherent collective motion ("flocking") at a solid-liquid interface. The polar order parameter and concentration of a collection of "active" (self-propelled) particles at a planar interface between a…

Soft Condensed Matter · Physics 2022-01-05 Niladri Sarkar , Abhik Basu , John Toner

Flocking, as paradigmatically exemplified by birds, is the coherent collective motion of active agents. As originally conceived, flocking emerges through alignment interactions between the agents. Here, we report that flocking can also…

Soft Condensed Matter · Physics 2024-07-16 Suchismita Das , Matteo Ciarchi , Ziqi Zhou , Jing Yan , Jie Zhang , Ricard Alert

We develop the hydrodynamic theory of dry, polar ordered, active matter (``flocking") with autochemotaxis; i.e., self-propelled entities moving in the same direction, each emitting a substance which attracts the others (e.g., ants). We find…

Soft Condensed Matter · Physics 2024-01-22 Maxx Miller , John Toner

We present a hydrodynamic model of flocking that generalizes the familiar Toner-Tu equations to incorporate turning inertia of well-polarized flocks. The continuum equations controlled by only two dimensionless parameters, orientational…

Soft Condensed Matter · Physics 2016-01-20 Xingbo Yang , M. Cristina Marchetti

Bacteria commonly live in structured communities that affect human health and influence ecological systems. Heterogeneous populations, such as motile and non-motile populations, often coexist in bacteria communities. Motile subpopulations…

Biological Physics · Physics 2019-06-19 Haoran Xu , Justas Dauparas , Debasish Das , Eric Lauga , Yilin Wu

Many motile biological cells navigate along concentration gradients of signaling molecules: This chemotaxis guides for instance sperm cells from marine invertebrates, which have to find egg cells in the ocean. While chemotaxis has been…

Biological Physics · Physics 2022-09-21 Steffen Lange , Benjamin M. Friedrich

We investigate the collective behavior of sterically interacting self-propelled particles confined in a harmonic potential. Our theoretical and numerical study unveils the emergence of distinctive collective polar organizations, revealing…

Soft Condensed Matter · Physics 2024-05-29 Daniel Canavello , Rubens H. Damascena , Leonardo R. E. Cabral , Clécio C. de Souza Silva

We investigate the collective dynamics of self-propelled particles able to probe and anticipate the orientation of their neighbors. We show that a simple anticipation strategy hinders the emergence of homogeneous flocking patterns. Yet,…

Statistical Mechanics · Physics 2015-06-23 Alexandre Morin , Jean-Baptiste Caussin , Christophe Eloy , Denis Bartolo