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In addition to enabling movement towards environments with favourable living conditions, swimming by microorganisms has also been linked to enhanced mixing and improved nutrient uptake by their populations. Experimental studies have shown…

Fluid Dynamics · Physics 2017-11-07 Blaise Delmotte , Eric E Keaveny , Eric Climent , Franck Plouraboué

We study the diffusion of a Brownian probe particle of size $R$ in a dilute dispersion of active Brownian particles (ABPs) of size $a$, characteristic swim speed $U_0$, reorientation time $\tau_R$, and mechanical energy $k_s T_s = \zeta_a…

Soft Condensed Matter · Physics 2017-05-17 Eric W. Burkholder , John F. Brady

Non-reciprocal interactions play a key role in shaping transport in active and passive systems, giving rise to striking nonequilibrium behavior. Here, we study the dynamics of a tracer -- active or passive -- embedded in a bath of active or…

Statistical Mechanics · Physics 2026-01-08 Subhajit Paul , Debasish Chaudhuri

Brownian motion is essential for describing diffusion in systems ranging from simple to complex liquids. Unlike simple liquids, which consist of only a solvent, complex liquids, such as colloidal suspensions or the cytoplasm of a cell, are…

Soft Condensed Matter · Physics 2025-02-27 Jeffrey C. Everts , Robert Hołyst , Karol Makuch

The induced diffusion of tracers in a bacterial suspension is studied theoretically and experimentally at low bacterial concentrations. Considering the swimmer-tracer hydrodynamic interactions at low-Reynolds number and using a kinetic…

Soft Condensed Matter · Physics 2015-06-11 G. L. Miño , J. Dunstan , A. Rousselet , E. Clement , R. Soto

We study the dynamics of a tracer in a dense mixture of particles connected to different thermostats. Starting from the overdamped Langevin equations that describe the evolution of the system, we derive the expression of the self-diffusion…

Soft Condensed Matter · Physics 2022-12-28 Marie Jardat , Vincent Dahirel , Pierre Illien

Swimming bacteria create long-range velocity fields that stir a large volume of fluid and move around passive particles dispersed in the fluid. Recent experiments and simulations have shown that long-time mean-squared displacement of…

Soft Condensed Matter · Physics 2013-08-20 Alexander Morozov , Davide Marenduzzo

Brownian motion is widely used as a paradigmatic model of diffusion in equilibrium media throughout the physical, chemical, and biological sciences. However, many real world systems, particularly biological ones, are intrinsically…

Statistical Mechanics · Physics 2020-06-04 Kiyoshi Kanazawa , Tomohiko G. Sano , Andrea Cairoli , Adrian Baule

Understanding particle motion in narrow channels is essential to guide progress in numerous applications, from filtration to vascular transport. Thermal or active fluctuations of channel walls for fluid-filled channels can slow down or…

Soft Condensed Matter · Physics 2023-12-29 Yating Wang , David S. Dean , Sophie Marbach , Ruben Zakine

Ensembles of biological and artificial microswimmers produce long-range velocity fields with strong nonequilibrium fluctuations, which result in dramatic increase diffusivity of embedded particles (tracers). While such enhanced diffusivity…

Soft Condensed Matter · Physics 2019-02-20 Sergey Belan , Mehran Kardar

Swimming microorganisms rely on effective mixing strategies to achieve efficient nutrient influx. Recent experiments, probing the mixing capability of unicellular biflagellates, revealed that passive tracer particles exhibit anomalous…

Soft Condensed Matter · Physics 2013-02-04 Irwin M. Zaid , Jörn Dunkel , Julia M. Yeomans

We consider the tracer diffusion $D_{rr}$ that arises from the run-and-tumble motion of low Reynolds number swimmers, such as bacteria. Assuming a dilute suspension, where the bacteria move in uncorrelated runs of length $\lambda$, we…

Soft Condensed Matter · Physics 2015-06-16 Dmitri O. Pushkin , Julia M. Yeomans

We consider two systems of active swimmers moving close to a solid surface, one being a living population of wild-type \textit{E. coli} and the other being an assembly of self-propelled Au-Pt rods. In both situations, we have identified two…

Populations of swimming microorganisms produce fluid motions that lead to dramatically enhanced diffusion of tracer particles. Using simulations of suspensions of swimming particles in a periodic domain, we capture this effect and show that…

Soft Condensed Matter · Physics 2008-07-22 Patrick T. Underhill , Juan P. Hernández-Ortiz , Michael D. Graham

The dynamics of a tracer molecule near a fluid membrane is investigated, with particular emphasis given to the interplay between the instantaneous position of the particle and membrane fluctuations. It is found that hydrodynamic…

Soft Condensed Matter · Physics 2009-11-11 Thomas Bickel

Tracer particles immersed in suspensions of biological microswimmers such as E. coli or C. reinhardtii display phenomena unseen in conventional equilibrium systems, including strongly enhanced diffusivity relative to the Brownian value and…

Soft Condensed Matter · Physics 2022-01-27 Henrik Nordanger , Alexander Morozov , Joakim Stenhammar

In this fluid dynamics video, we demonstrate the microscale mixing enhancement of passive tracer particles in suspensions of swimming microalgae, Chlamydomonas reinhardtii. These biflagellated, single-celled eukaryotes (10 micron diameter)…

We investigate the dynamics of a massive tracer particle coupled to an interacting active bath, modeled as a harmonic chain of overdamped active particles analytically, with an aim to understand the impact of bath interactions and activity…

Statistical Mechanics · Physics 2025-09-03 Ritwick Sarkar , Ion Santra

Active matter exhibits various forms of non-equilibrium states in the absence of external forcing, including macroscopic steady-state currents. Such states are often too complex to be modelled from first principles and our understanding of…

Soft Condensed Matter · Physics 2020-09-17 Viktor Škultéty , Cesare Nardini , Joakim Stenhammar , Davide Marenduzzo , Alexander Morozov

A self-propelled artificial microswimmer is often modeled as a ballistic Brownian particle moving with constant speed aligned along one of its axis, but changing direction due to random collisions with the environment. Similarly to thermal…

Soft Condensed Matter · Physics 2015-12-08 Pulak K. Ghosh , Yunyun Li , Giampiero Marchegiani , Fabio Marchesoni
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