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Understanding swimming in soft yielding media is challenging due to their complex deformation response to the swimmer's motion. We experimentally show that a scallop-inspired swimmer with reciprocally flapping wings generates locomotion in…

Soft Condensed Matter · Physics 2024-12-09 Hongyi Xiao , Harol Torres , Achim Sack , Thorsten Pöschel

Living systems maintain or increase local order by working against the Second Law of Thermodynamics. Thermodynamic consistency is restored as they dissipate heat, thereby increasing the net entropy of their environment. Recently introduced…

Biological Physics · Physics 2021-05-03 Dominic J. Skinner , Jörn Dunkel

Active emulsions can spontaneously form self-propelled droplets or phoretic micropumps. It has been predicted that the interaction with their self-generated chemical fields can lead to multistable higher-order flows and chemodynamic…

From bacteria and sperm cells to artificial microrobots, self-propelled microscopic objects at low Reynolds numbers often perceive fluctuating mechanical and chemical stimuli and contact exterior wall boundaries both in nature and the…

Soft Condensed Matter · Physics 2023-12-22 Yoshiki Hiruta , Kenta Ishimoto

We formulate and solve the equations governing the dynamics of a microscopic artificial swimmer composed of a head and of a tail made of a thin film of permanent magnetic material. This is a variant of the model swimmer proposed by Dreyfus…

Soft Condensed Matter · Physics 2017-02-15 Francois Alouges , Antonio DeSimone , Laetitia Giraldi , Marta Zoppello

The unique properties of suspensions containing both active (self-propelling) and passive matter, arising from the nonequilibrium nature of these systems, have been widely studied (e.g., enhanced diffusion, phase separation, and directed…

Computational Physics · Physics 2019-10-23 Ryan Krafnic , Angel E. Garcia

Swimming consists by definition in propelling through a fluid by means of bodily movements. Thus, from a mathematical point of view, swimming turns into a control problem for which the controls are the deformations of the swimmer. The aim…

Optimization and Control · Mathematics 2017-02-15 Thomas Chambrion , Laetitia Giraldi , Alexandre Munnier

Understanding how microscopic motility shapes emergent collective behaviors is a challenging task in active matter, especially when self-propulsion is regulated by external cues or via quorum-sensing interactions. To address this problem,…

Statistical Mechanics · Physics 2026-04-13 Alberto Dinelli , Pietro Luigi Muzzeddu

In biological systems, microswimmers often propel themselves through complex media. However, many aspects of swimming mechanisms in non-Newtonian fluids remain unclear. This study considers the propulsion of two types of single spherical…

Fluid Dynamics · Physics 2024-10-14 Takuya Kobayashi , Ryoichi Yamamoto

In analogy to nanoscopic molecules that are composed of individual atoms, we consider an active "microswimmer molecule". It is built up from three individual magnetic colloidal microswimmers that are connected by harmonic springs and…

Soft Condensed Matter · Physics 2016-04-12 Sonja Babel , Hartmut Löwen , Andreas M. Menzel

We numerically investigate the motion of active artificial microswimmers diffusing in a fuel concentration gradient. We observe that, in the steady state, their probability density accumulates in the low-concentration regions, whereas a…

Soft Condensed Matter · Physics 2025-08-18 Pulak K. Ghosh , Yunyun Li , Fabio Marchesoni , Franco Nori

A liquid droplet, immersed into a Newtonian fluid, can be propelled solely by internal flow. In a simple model, this flow is generated by a collection of point forces, which represent externally actuated devices or model autonomous…

Fluid Dynamics · Physics 2021-03-10 Reiner Kree , Leon Rueckert , Annette Zippelius

Nonequilibrium processes break time-reversal symmetry and generate entropy. Living systems are driven out-of-equilibrium at the microscopic level of molecular motors that exploit chemical potential gradients to transduce free energy to…

Statistical Mechanics · Physics 2022-12-06 Eden Nitzan , Aishani Ghosal , Gili Bisker

In nature, many unicellular organisms are able to swim with the help of beating filaments, where local energy input leads to cooperative undulatory beating motion. Here, we investigate by employing reinforcement learning how undulatory…

Biological Physics · Physics 2025-11-04 Fumiya Tokoro , Hideki Takayama , Shinji Deguchi , Andreas Zöttl , Daiki Matsunaga

Inviscid computational results are presented on a self-propelled swimmer modeled as a virtual body combined with a two-dimensional hydrofoil pitching intermittently about its leading edge. Lighthill (1971) originally proposed that this…

Fluid Dynamics · Physics 2018-02-14 Emre Akoz , Keith W. Moored

Synthetic microswimmers mimicking biological movements at the microscale have been developed in recent years. Actuating helical magnetic materials with a homogeneous rotating magnetic field is one of the most widespread techniques for…

Soft Condensed Matter · Physics 2019-03-27 Felix Bachmann Klaas Bente , Agnese Codutti , Damien Faivre

Transport of microscopic objects across biological membranes usually involves membrane deformation to enclose the object followed by detachment of the engulfed particle. However, in artificial membranes, this last topological remodelling…

Soft Condensed Matter · Physics 2025-03-25 Florent Fessler , Pierre Muller , Antonio Stocco

Time-reversal symmetry breaking and entropy production are universal features of nonequilibrium phenomena. Despite its importance in the physics of active and living systems, the entropy production of systems with many degrees of freedom…

We report measurements of global dissipated power within a turbulent flow homogeneously forced at small scale by a new forcing technique. The forcing is random in both time and space within the fluid by using magnetic particles in an…

Fluid Dynamics · Physics 2017-10-31 Eric Falcon , Jean-Claude Bacri , Claude Laroche

Entropy production rate (EPR) is often effective to describe how a structure is self-organized in a nonequilibrium thermodynamic system. The "minimum EPR principle" is widely applicable to characterizing self-organized structures, but is…

Plasma Physics · Physics 2015-11-05 Yohei Kawazura , Zensho Yoshida