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Related papers: Wrinkling instability in unsupported epithelial sh…

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In cell extrusion, a cell embedded in an epithelial monolayer loses its apical or basal surface and is subsequently squeezed out of the monolayer by neighboring cells. Cell extrusions occur during apoptosis, epithelial-mesenchymal…

Cell Behavior · Quantitative Biology 2020-04-24 Satoru Okuda , Koichi Fujimoto

Numerical simulations of thin sheets undergoing large deformations are computationally challenging. Depending on the scenario, they may spontaneously buckle, wrinkle, fold, or crumple. Nature's thin tissues often experience significant…

Soft Condensed Matter · Physics 2015-08-03 Roman Vetter , Norbert Stoop , Falk K. Wittel , Hans J. Herrmann

We consider a cellular monolayer, described using a vertex-based model, for which cells form a spatially disordered array of convex polygons that tile the plane. Equilibrium cell configurations are assumed to minimize a global energy…

Cell Behavior · Quantitative Biology 2018-05-30 Alexander Nestor-Bergmann , Emma Johns , Sarah Woolner , Oliver E. Jensen

Non-Euclidean plates are thin elastic bodies having no stress-free configuration, hence exhibiting residual stresses in the absence of external constraints. These bodies are endowed with a three-dimensional reference metric, which may not…

Soft Condensed Matter · Physics 2009-05-29 Efi Efrati , Eran Sharon , Raz Kupferman

An analytical model based on variational principles for a thin-walled stiffened plate subjected to axial compression is presented. A system of nonlinear differential and integral equations is derived and solved using numerical continuation.…

Materials Science · Physics 2014-06-10 M. Ahmer Wadee , Maryam Farsi

Predicting the large-amplitude deformations of thin elastic sheets is difficult due to the complications of self-contact, geometric nonlinearities, and a multitude of low-lying energy states. We study a simple two-dimensional setting where…

Biological processes, from morphogenesis to tumor invasion, spontaneously generate shear stresses inside living tissue. The mechanisms that govern the transmission of mechanical forces in epithelia and the collective response of the tissue…

Soft Condensed Matter · Physics 2022-11-29 Junxiang Huang , James O. Cochran , Suzanne M. Fielding , M. Cristina Marchetti , Dapeng Bi

The modeling of the elastic properties of disordered or nanoscale solids requires the foundations of the theory of elasticity to be revisited, as one explores scales at which this theory may no longer hold. The only cases for which…

Materials Science · Physics 2007-05-23 I. Goldhirsch , C. Goldenberg

Buckling instabilities driven by tissue growth underpin key developmental events such as the folding of the brain. Tissue growth is disordered due to cell-to-cell variability, but the effects of this variability on buckling are unknown.…

Soft Condensed Matter · Physics 2025-12-12 Rahul G. Ramachandran , Ricard Alert , Pierre A. Haas

Twisted and rope-like assemblies of filamentous molecules are common and vital structural elements in cells and tissue of living organisms. We study the intrinsic frustration occurring in these materials between the two-dimensional…

Soft Condensed Matter · Physics 2015-06-03 Amir Azadi , Gregory M. Grason

We show that thin sheets under boundary confinement spontaneously generate a universal self-similar hierarchy of wrinkles. From simple geometry arguments and energy scalings, we develop a formalism based on wrinklons, the transition zone in…

We propose two toy-models to describe, predict, and interpret the wrinkles appearing on the surface of skin when it is sheared. With the first model, we account for the lines of greatest tension present in human skin by subjecting a layer…

Soft Condensed Matter · Physics 2013-06-04 Pasquale Ciarletta , Michel Destrade , Artur L. Gower

We study the dynamic coarsening of wrinkles in an elastic sheet that is compressed while lying on a thin layer of viscous liquid. When the ends of the sheet are instantaneously brought together by a small distance, viscous resistance…

Fluid Dynamics · Physics 2017-03-24 Ousmane Kodio , Ian M. Griffiths , Dominic Vella

Thin dielectric elastomers with compliant electrodes exhibit various types of instability under the action of electromechanical loading. Guided by the thermodynamically-based formulation of Fosdick and Tang (J. Elasticity 88, 255-297,…

Soft Condensed Matter · Physics 2019-08-20 G. Zurlo

A slender-thread model is derived to explore the Rayleigh-Plateau instability of a filament of elasto-viscoplastic fluid. Without elasticity, a finite yield stress suppresses any linear instability for a filament of constant radius.…

Fluid Dynamics · Physics 2025-12-25 James D. Shemilt , Neil J. Balmforth

Thin elastic sheets appear in systems ranging from graphene to biological membranes, where phenomena such as wrinkling, folding, and thermal fluctuations originate from geometric nonlinearities. These effects are treated within weakly…

Soft Condensed Matter · Physics 2025-12-04 Yael Cohen , Animesh Pandey , Yafei Zhang , Cy Maor , Michael Moshe

Treating the epithelium as an incompressible fluid adjacent to a viscoelastic stroma, we find a novel hydrodynamic instability that leads to the formation of protrusions of the epithelium into the stroma. This instability is a candidate for…

Biological Physics · Physics 2012-03-09 Markus Basan , Jean-Francois Joanny , Jacques Prost , Thomas Risler

Recent experiments have shown that surface stresses in soft materials can have a significant strain-dependence. Here we explore the implications of this surface elasticity to show how, and when, we expect it to arise. We develop the…

Soft Condensed Matter · Physics 2020-05-28 Robert W. Style , Qin Xu

We develop a general incremental framework for hyperelastic solids whose surfaces exhibit both stretch-dependent and curvature-dependent elastic behavior. Building upon a variational formulation of curvature-dependent surface elasticity, we…

Mathematical Physics · Physics 2026-01-08 Xiang Yu , Michal Šmejkal , Martin Horák

A controlled surface wrinkling pattern has been widely used in diverse applications, such as stretchable electronics, smart windows, and haptics. Here, we focus on hexagonal wrinkling patterns because of their great potentials in realizing…

Soft Condensed Matter · Physics 2021-02-24 Teng Zhang
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