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Tissue fluidity regulates many critical biological processes, including embryonic development, wound healing, and cancer metastasis. In confluent epithelia, where cell packing fraction is effectively fixed, the prevailing paradigm…

Biological Physics · Physics 2026-03-09 Pradip K. Bera , Anh Q. Nguyen , Molly McCord , Dapeng Bi , Jacob Notbohm

We study theoretically the shapes of a dividing epithelial monolayer of cells lying on top of an elastic stroma. The negative tension created by cell division provokes a buckling instability at a finite wave vector leading to the formation…

Biological Physics · Physics 2012-03-12 Edouard Hannezo , Jacques Prost , Jean-Francois Joanny

Wrinkling instabilities of thin elastic sheets can be used to generate periodic structures over a wide range of length scales. Viscosity of the thin elastic sheet or its surrounding medium has been shown to be responsible for dynamic…

Soft Condensed Matter · Physics 2024-01-25 Veit Krause , Axel Voigt

Recent experiments by Kantsler et. al. (2007) have shown that the relaxational dynamics of a vesicle in external elongation flow is accompanied by the formation of wrinkles on a membrane. Motivated by these experiments we present a theory…

Soft Condensed Matter · Physics 2013-05-29 K. S. Turitsyn , S. S. Vergeles

The formation of periodic wrinkles in soft layered materials due to mechanical instabilities is prevalent in nature and has been proposed for use in multiple applications. However, such phenomena have been explored predominantly in…

Non-Euclidean plates are a subset of the class of elastic bodies having no stress-free configuration. Such bodies exhibit residual stress when relaxed from all external constraints, and may assume complicated equilibrium shapes even in the…

Soft Condensed Matter · Physics 2009-11-13 Efi Efrati , Eran Sharon , Raz Kupferman

Nematic elastomers are rubbery solids which have liquid crystals incorporated into their polymer chains. These materials display many unusual mechanical properties, one such being the ability to form fine-scale microstructure. In this work,…

Soft Condensed Matter · Physics 2017-04-05 Paul Plucinsky , Kaushik Bhattacharya

Motivated by the redrawing of hot glass into thin sheets, we investigate the shape and stability of a thin viscous sheet that is inhomogeneously stretched in an imposed non-uniform temperature field. We first determine the associated base…

Fluid Dynamics · Physics 2017-07-28 Siddarth Srinivasan , Zhiyan Wei , L. Mahadevan

Wrinkling of stretched elastic sheets is widely observed, and the scaling relations between the amplitude and wavelength of the wrinkles have been proposed by Cerda and Mahadevan. However, the surface effects should be taken into account…

Mesoscale and Nanoscale Physics · Physics 2013-11-22 Jie Gu

Surface growth is a crucial component of many natural and artificial processes from cell proliferation to additive manufacturing. In elastic systems surface growth is usually accompanied by the development of geometrical incompatibility…

Soft Condensed Matter · Physics 2019-05-08 Lev Truskinovsky , Giuseppe Zurlo

A thin flat rectangular plate supported on its edges and subjected to in-plane loading exhibits stable post-buckling behaviour. However, the introduction of a nonlinear (softening) elastic foundation may cause the response to become…

Pattern Formation and Solitons · Physics 2016-03-18 M. Khurram Wadee , David J. B. Lloyd , Andrew P. Bassom

A thin, elastic sheet floating on the surface of a liquid bath wrinkles when poked at its centre. We study the onset of wrinkling as well as the evolution of the pattern as indentation progresses far beyond the wrinkling threshold. We use…

Soft Condensed Matter · Physics 2018-07-31 Dominic Vella , Benny Davidovitch

Epithelial monolayers are a central building block of complex organisms. Topological defects have emerged as important elements for single cell behavior in flat epithelia. Here we theoretically study such defects in a three-dimensional…

Biological Physics · Physics 2025-02-19 Oliver M. Drozdowski , Ulrich S. Schwarz

We consider a disk-shaped thin elastic sheet bonded to a compliant sphere. (Our sheet can slip along the sphere; the bonding controls only its normal displacement.) If the bonding is stiff (but not too stiff), the geometry of the sphere…

Analysis of PDEs · Mathematics 2017-04-12 Peter Bella , Robert V. Kohn

Thin elastic sheets and membranes are known to wrinkle when they are stretched -- the associated physics is highly non-linear. The mechanics of thin films that exhibit unusual behavior upon stretching, when they possess auxetic structure,…

Soft Condensed Matter · Physics 2020-07-22 Alessandra Bonfanti , Atul Bhaskar

Wrinkling is the phenomenon of out-of-plane deformation patterns in thin walled structures, as a result of a local compressive (internal) loads in combination with a large membrane stiffness and a small but non-zero bending stiffness.…

Numerical Analysis · Mathematics 2025-03-20 H. M. Verhelst , M. Möller , J. H. Den Besten

A ubiquitous motif in nature is the self-similar hierarchical buckling of a thin lamina near its margins. This is seen in leaves, flowers, fungi, corals, and marine invertebrates. We investigate this morphology from the perspective of…

We investigate the wrinkling dynamics of an elastic filament immersed in a viscous fluid submitted to compression at a finite rate with experiments and by combining geometric nonlinearities, elasticity, and slender body theory. The drag…

Soft Condensed Matter · Physics 2017-08-30 Julien Chopin , Moumita Dasgupta , Arshad Kudrolli

We present a hydrodynamic model of spreading epithelial monolayers as polar viscous fluids, with active contractility and traction on the substrate. The combination of both active forces generate an instability that leads to nonlinear…

Soft Condensed Matter · Physics 2017-05-02 C. Blanch-Mercader , J. Casademunt

We show that a viscoelastic thin sheet driven out of equilibrium by active structural remodelling develops a rich variety of shapes as a result of a competition between viscous relaxation and activity. In the regime where active processes…

Soft Condensed Matter · Physics 2020-02-26 D. A. Matoz-Fernandez , Fordyce A. Davidson , Nicola R. Stanley-Wall , Rastko Sknepnek