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Fish swim with flexible fins that stand in stark contrast to the rigid propulsors of engineered vehicles. Using numerical simulations of the dynamics of flow-structure interaction, we have found that dorso-ventral deformation in flexible…

Fluid Dynamics · Physics 2026-04-16 Sushrut Kumar , Matthew J. McHenry , Jung-Hee Seo , Rajat Mittal

In isotropic fluids like water, micrometer-scale swimmers have evolved swim strokes to translate despite their tiny size. As described by Purcell in his Scallop Theorem, reciprocal motions, like those performed by a scallop, cannot drive…

Active colloidal microswimmers serve as archetypical active fluid systems, and as models for biological swimmers. Here, by studying in detail their velocity traces, we find robust power-law intermittency with system-dependent exponential…

Soft Condensed Matter · Physics 2023-10-27 Nick Oikonomeas-Koppasis , Stefania Ketzetzi , Daniela J. Kraft , Peter Schall

Microswimmers (planktonic microorganisms or artificial active particles) immersed in a fluid interact with the ambient flow, altering their trajectories. In surface gravity waves, a common goal for microswimmers is vertical migration (e.g.,…

Soft Condensed Matter · Physics 2022-03-01 Kunlin Ma , Nimish Pujara , Jean-Luc Thiffeault

The motility of microorganisms is influenced greatly by their hydrodynamic interactions with the fluidic environment they inhabit. We show by direct experimental observation of the bi-flagellated alga Chlamydomonas reinhardtii that fluid…

Biological Physics · Physics 2015-11-03 Boyang Qin , Arvind Gopinath , Jing Yang , Jerry P Gollub , Paulo E Arratia

The swimming trajectories of self-propelled organisms or synthetic devices in a viscous fluid can be altered by hydrodynamic interactions with nearby boundaries. We explore a multipole description of swimming bodies and provide a general…

Fluid Dynamics · Physics 2012-11-30 Saverio E. Spagnolie , Eric Lauga

Centipede-like robots offer unique locomotion advantages due to their small cross-sectional area for accessing confined spaces, and their redundant legs enhance robustness in cluttered environments such as search-and-rescue and pipe…

Elongate animals and robots use undulatory body waves to locomote through diverse environments. Geometric mechanics provides a framework to model and optimize such systems in highly damped environments, connecting a prescribed shape change…

Robotics · Computer Science 2025-10-21 Jianfeng Lin , Tianyu Wang , Baxi Chong , Matthew Fernandez , Zhaochen Xu , Daniel I. Goldman

Flagellar-driven locomotion plays a critical role in bacterial attachment and colonization of surfaces, contributing to the risks of contamination and infection. Tremendous attempts to uncover the underlying principles governing bacterial…

Soft Condensed Matter · Physics 2025-02-19 Xin-Xin Xu , Yangguang Tian , Yuhe Pu , Bingchen Che , Hao Luo , Yanan Liu , Yan-Jun Liu , Guangyin Jing

In this communication we address some hydrodynamic aspects of recently revisited drift mechanism of biogenic mixing in the ocean (Katija and Dabiri, Nature vol. 460, pp. 624-626, 2009). The relevance of the locomotion gait at various…

Fluid Dynamics · Physics 2015-05-19 A. M. Leshansky , L. M. Pismen

Animals achieve robust locomotion by offloading regulation from the brain to physical couplings within the body. In contrast, locomotion in artificial systems often depends on centralized processors. We introduce a rapid and autonomous…

Soft Condensed Matter · Physics 2025-05-16 Alberto Comoretto , Harmannus A. H. Schomaker , Johannes T. B. Overvelde

We consider a finite-dimensional model for the motion of microscopic organisms whose propulsion exploits the action of a layer of cilia covering its surface. The model couples Newton's laws driving the organism, considered as a rigid body,…

Optimization and Control · Mathematics 2009-09-29 Mario Sigalotti , Jean-Claude Vivalda

Swimming micro-organisms such as flagellated bacteria and sperm cells have fascinating locomotion capabilities. Inspired by their natural motion, there is an ongoing effort to develop artificial robotic nano-swimmers for potential in-body…

Fluid Dynamics · Physics 2023-05-31 Jithu Paul , Yizhar Or , Oleg Gendelman

The survival of many microorganisms, like \textit{Leptospira} or \textit{Spiroplasma} bacteria, can depend on their ability to navigate towards regions of favorable viscosity. While this ability, called viscotaxis, has been observed in…

Soft Condensed Matter · Physics 2018-05-23 Benno Liebchen , Paul Monderkamp , Borge ten Hagen , Hartmut Löwen

Active particles, which interact hydrodynamically, display a remarkable variety of emergent collective phenomena. We use squirmers to model spherical microswimmers and explore the collective behavior of thousands of them under the influence…

Fluid Dynamics · Physics 2017-10-31 Jan-Timm Kuhr , Johannes Blaschke , Felix Rühle , Holger Stark

The motion of a rotating helical body in a viscoelastic fluid is considered. In the case of force-free swimming, the introduction of viscoelasticity can either enhance or retard the swimming speed and locomotive efficiency, depending on the…

Biological Physics · Physics 2013-08-15 Saverio E. Spagnolie , Bin Liu , Thomas R. Powers

Active diffusiophoresis - swimming through interaction with a self-generated, neutral, solute gradient - is a paradigm for autonomous motion at the micrometer scale. We study this propulsion mechanism within a linear response theory.…

Statistical Mechanics · Physics 2015-05-30 Benedikt Sabass , Udo Seifert

We demonstrate with experiments and simulations how microscopic self-propelled particles navigate through environments presenting complex spatial features, which mimic the conditions inside cells, living organisms and future lab-on-a-chip…

Biological Physics · Physics 2011-09-22 Giovanni Volpe , Ivo Buttinoni , Dominik Vogt , Hans-Juergen Kuemmerer , Clemens Bechinger

We use particle simulations to reveal two distinct propulsion mechanisms for a scallop-like swimmer to locomote itself in granular media by reciprocally flapping its wings. Based on the discrete element method, we examine the kinematics and…

Soft Condensed Matter · Physics 2025-10-28 Amir Nazemi , Hongyi Xiao

The possibility of microscopic swimming by extraction of energy from an external flow is discussed, focusing on the migration of a simple trimer across a linear shear flow. The geometric properties of swimming, together with the possible…

Soft Condensed Matter · Physics 2015-05-18 Piero Olla