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We investigate the emergence of global alignment in colonies of dividing rod-shaped cells under confinement. Using molecular dynamics simulations and continuous modeling, we demonstrate that geometrical anisotropies in the confining…

Soft Condensed Matter · Physics 2021-01-26 Zhihong You , Daniel J. G. Pearce , Luca Giomi

Non-motile elongated bacteria confined in two-dimensional open micro-channels can exhibit collective motion and form dense monolayers with nematic order if the cells proliferate, i.e., grow and divide. Using soft molecular dynamics…

Soft Condensed Matter · Physics 2013-07-25 Sirio Orozco-Fuentes , Denis Boyer

In densely-packed two-dimensional systems of growing cells, such as rod-shaped bacteria, a number of experimental and numerical studies report distinct patterns of nematic orientational order in the presence of confinement. So far, these…

Soft Condensed Matter · Physics 2025-06-13 Lukas Hupe , Jonas Isensee , Ramin Golestanian , Philip Bittihn

Living active collectives have evolved with remarkable self-patterning ability to meet the physical and biological constraints for growth and survival. However, how complex multicellular patterns emerge from a single founder cell remains…

Biological Physics · Physics 2022-11-30 Changhao Li , Japinder Nijjer , Luyi Feng , Qiuting Zhang , Jing Yan , Sulin Zhang

Motivated by recent experiments on growing fibroblasts, we examine the development of nematic order in a colony of elongated cells proliferating on a nematic elastomer substrate. After sparse seeding, the cells divide and grow into locally…

Soft Condensed Matter · Physics 2026-01-06 Toshi Parmar , Fridtjof Brauns , Yimin Luo , M. Cristina Marchetti

Collective dynamics in proliferating anisotropic particle systems arise from an interplay between growth, division, and mechanical interactions, often mediated by particle shape. In classical models of prolate, rod-like growth, flow-induced…

Soft Condensed Matter · Physics 2025-12-02 Jonas Isensee , Lukas Hupe , Philip Bittihn

The ordering of rod-like particles in soft, deformable containers emerges from the interplay of anisotropic interactions, geometric confinement, and boundary compliance. This competition couples internal particle organization to container…

Soft Condensed Matter · Physics 2026-05-22 Stijn van der Ham , André F. V. Matias , Marjolein Dijkstra , Hanumantha Rao Vutukuri

Monolayers of growing bacteria, confined within channel geometries, exhibit self-organization into a highly aligned laminar state along the axis of the channel. Although this phenomenon has been observed in experiments and simulations under…

Soft Condensed Matter · Physics 2024-11-04 Blake Langeslay , Will Fahy , Gabriel Juarez

Directed growth, anisotropic cell shapes, and confinement drive self-organization in multicellular systems. We investigate the influence of particle shape on the distribution and dynamics of nematic microdomains in a minimal in-silico model…

Soft Condensed Matter · Physics 2025-12-03 Jonas Isensee , Philip Bittihn

The ability of cells to reorganize in response to external stimuli is important in areas ranging from morphogenesis to tissue engineering. Elongated cells can co-align due to steric effects, forming states with local order. We show that…

Growing monolayers of rod-shaped bacteria exhibit local alignment similar to extensile active nematics. When confined in a channel or growing inward from a ring, the local nematic order of these monolayers changes to a global ordering with…

Soft Condensed Matter · Physics 2024-11-04 Blake Langeslay , Gabriel Juarez

Mechanical interactions among cells in a growing microbial colony can significantly influence the colony's spatial genetic structure and, thus, evolutionary outcomes such as the fates of rare mutations. Here, we computationally investigate…

Soft Condensed Matter · Physics 2025-01-22 Mateusz Ratman , Jimmy Gonzalez Nuñez , Daniel A. Beller

Vertically vibrated rod-shaped granular materials confined to quasi-2D containers self organize into distinct patterns. We find, consistent with theory and simulation, a density dependent isotropic-nematic transition. Along the walls, rods…

Soft Condensed Matter · Physics 2009-11-11 J. Galanis , D. Harries , D. L. Sackett , W. Losert , R. Nossal

Bacterial colonies growing on surfaces are shaped by mechanical stresses transmitted through the community, governed by the balance between cell growth and steric and cell-substrate interactions. Using overdamped dynamics simulations of…

Soft Condensed Matter · Physics 2026-01-14 Samaneh Rahbar , Ludger Santen , Reza Shaebani

Recent studies have shown that packings of cells, both eukaryotic cellular tissues and growing or swarming bacterial colonies, can often be understood as active nematic fluids. A key property of volume-conserving active nematic model…

Soft Condensed Matter · Physics 2022-05-12 Fabian Jan Schwarzendahl , Daniel A. Beller

Growing tissue and bacterial colonies are active matter systems where cell divisions and cellular motion generate active stress. Although they operate in the non-equilibrium regime, these biological systems can form large-scale ordered…

Biological Physics · Physics 2019-05-28 Yusuf Ilker Yaman , Esin Demir , Roman Vetter , Askin Kocabas

We report on experiments to measure the temporal and spatial evolution of packing arrangements of anisotropic, cylindrical granular material, using high-resolution capacitive monitoring. In these experiments, the particle configurations…

Materials Science · Physics 2009-10-31 Fernando X. Villarruel , Benjamin E. Lauderdale , Daniel M. Mueth , Henrich M. Jaeger

We investigate the effect of an anisotropic substrate on the turbulent dynamics of a simulated two dimensional active nematic. This is introduced as an anisotropic friction and an effective anisotropic viscosity, with the orientation of the…

Soft Condensed Matter · Physics 2019-06-12 D. J. G. Pearce

Monolayers of growing non-motile rod-shaped bacteria act as active nematic materials composed of hard particles rather than the flexible components of other commonly studied active nematics. The organization of these granular monolayers has…

Soft Condensed Matter · Physics 2023-05-12 Blake Langeslay , Gabriel Juarez

Cell alignment often forms nematic order, which can lead to anomalous collective cell flow due to the so-called active force. Although it is appreciated that cell migration is driven by traction force, a quantitative evaluation of the…

Soft Condensed Matter · Physics 2024-02-27 Masahito Uwamichi , He Li , Zihui Zhao , Yisong Yao , Hideo Higuchi , Kyogo Kawaguchi , Masaki Sano
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