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We present a general framework for modeling a wide selection of flocking scenarios under free boundary conditions. Several variants have been considered - including examples for the widely observed behavior of hierarchically interacting…

Physics and Society · Physics 2019-04-23 Yongnan Jia , Tamas Vicsek

In this study, we investigate the phenomenon of collective motion in binary mixtures of self-propelled particles. We consider two particle species, each of which consisting of pointlike objects that propel with a velocity of constant…

Soft Condensed Matter · Physics 2013-11-20 Andreas M. Menzel

Designing microscopic and nanoscopic self-propelled particles and characterising their motion has become a major scientific challenge over the past decades. To this purpose, phoretic effects, namely propulsion mechanisms relying on local…

Soft Condensed Matter · Physics 2021-04-09 Pierre Illien , Ramin Golestanian , Ayusman Sen

A one-dimensional rule-based model for flocking, that combines velocity alignment and long-range centering interactions, is presented and studied. The induced cohesion in the collective motion of the self-propelled agents leads to a unique…

Statistical Mechanics · Physics 2015-01-19 V. Dossetti

We study the (hydro-)dynamics of multi-species driven by alignment. What distinguishes the different species is the protocol of their interaction with the rest of the crowd: the collective motion is described by different communication…

Analysis of PDEs · Mathematics 2020-10-20 Siming He , Eitan Tadmor

Numerical models indicate that collective animal behaviour may emerge from simple local rules of interaction among the individuals. However, very little is known about the nature of such interaction, so that models and theories mostly rely…

Populations and Evolution · Quantitative Biology 2009-11-13 M. Ballerini , N. Cabibbo , R. Candelier , A. Cavagna , E. Cisbani , I. Giardina , V. Lecomte , A. Orlandi , G. Parisi , A. Procaccini , M. Viale , V. Zdravkovic

One of the most striking phenomena in biological systems is the tendency for biological agents to spatially aggregate, and subsequently display further collective behaviours such as rotational motion. One prominent explanation for why…

Quantitative Methods · Quantitative Biology 2024-10-18 Thomas Stemler , Shannon Dee Algar , Jesse Zhou

The transition from a microscopic model for the movement of many particles to a macroscopic continuum model for a density flow is studied. The microscopic model for the free flow is completely deterministic, described by an interaction…

Statistical Mechanics · Physics 2021-01-12 Jennifer Weissen , Simone Göttlich , Dieter Armbruster

We investigate the emergence of cohesive flocking in open, boundless space using a multi-agent reinforcement learning framework. Agents integrate positional and orientational information from their closest topological neighbours and learn…

Soft Condensed Matter · Physics 2026-02-02 Martino Brambati , Antonio Celani , Marco Gherardi , Francesco Ginelli

Along a microtubule, certain active motors propel themselves in one direction whereas others propel themselves in the opposite direction. For example, the cargo transporting motor proteins dynein and kinesin propel themselves towards the…

Soft Condensed Matter · Physics 2016-04-01 Anton Souslov , Paul M. Goldbart

We review the observations and the basic laws describing the essential aspects of collective motion -- being one of the most common and spectacular manifestation of coordinated behavior. Our aim is to provide a balanced discussion of the…

Statistical Mechanics · Physics 2012-08-16 Tamás Vicsek , Anna Zafeiris

The collective motion of epithelial cells is a fundamental biological process which plays a significant role in embryogenesis, wound healing and tumor metastasis. While it has been broadly investigated for over a decade both in vivo and in…

Classical swarm models, exemplified by the Cucker--Smale framework, provide foundational insights into collective alignment but exhibit fundamental limitations in capturing the adaptive, heterogeneous behaviours intrinsic to living systems.…

Adaptation and Self-Organizing Systems · Physics 2025-09-08 Rene Fabregas , Jie Liao , Nisrine Outada

Many animal groups are heterogeneous and may even consist of individuals of different species, called mixed-species flocks. Mathematical and computational models of collective animal movement behaviour, however, typically assume that groups…

Adaptation and Self-Organizing Systems · Physics 2019-03-27 Gokul G. Nair , Athmanathan Senthilnathan , Srikanth K. Iyer , Vishwesha Guttal

We use topological data analysis and machine learning to study a seminal model of collective motion in biology [D'Orsogna et al., Phys. Rev. Lett. 96 (2006)]. This model describes agents interacting nonlinearly via attractive-repulsive…

Coherent collective motion is a widely observed phenomenon in active matter systems. Here, we report a flocking transition mechanism in a system of chemically interacting active colloidal particles sustained purely by chemo-repulsive…

Soft Condensed Matter · Physics 2025-11-04 Arvin Gopal Subramaniam , Sagarika Adhikary , Rajesh Singh

Using a minimal aggregation-based model, we address the efficient information transfer observed in natural flocks during collective turns. Specifically, we demonstrate that this feature can arise solely from the non-reciprocal nature of…

Adaptation and Self-Organizing Systems · Physics 2025-11-25 Joao Lizárraga , Marcus de Aguiar

We formulate a kinetic theory of two species of hard spheres undergoing reactive collisions that convert chemical energy into kinetic energy. The model describes an active species interacting with a passive background, labeled as birds and…

Statistical Mechanics · Physics 2026-01-07 Ruben Lier

Natural flocks (aligned) and swarms (non-aligned) both exhibit features of near-criticality, challenging their treatment as two ends of the same phase transition. We present a model for the aggregation of active individuals, in which their…

Adaptation and Self-Organizing Systems · Physics 2025-11-26 Joao Lizárraga , Marcus de Aguiar

Asymmetric obstacles can be exploited to direct the motion and induce sorting of run-and-tumbling particles. In this work, we show that flocking particles which follow the Vicsek model aligning rules experience a collective trapping in the…

Soft Condensed Matter · Physics 2019-12-02 Raul Martinez , Francisco Alarcon , Juan Luis Aragones , Chantal Valeriani