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Proliferation and motility are ubiquitous drivers of activity in biological systems. Here, we study a dense binary mixture of motile and proliferating particles with exclusively repulsive interactions, where homeostasis in the proliferating…

Soft Condensed Matter · Physics 2025-06-11 Lukas Hupe , Joanna M. Materska , David Zwicker , Ramin Golestanian , Bartlomiej Waclaw , Philip Bittihn

Biological active matter like the cytoskeleton or tissues are characterized by their ability to transform chemical energy into mechanical stress. In addition, it often exhibits orientational order, which is essential for many cellular and…

Biological Physics · Physics 2025-06-05 Ludovic Dumoulin , Carles Blanch-Mercader , Karsten Kruse

We study the universal behavior of a class of active colloids whose design is inspired by the collective dynamics of natural systems like schools of fish and flocks of birds. These colloids, with off-center repulsive interaction sites,…

Soft Condensed Matter · Physics 2024-10-10 Amir Nourhani

We introduce and theoretically investigate a minimal particle-based model for a new class of active matter where particles exhibit directional, volume-conserving division in confinement while interacting sterically, mimicking cells in early…

Soft Condensed Matter · Physics 2024-10-01 Samantha R. Lish , Lukas Hupe , Ramin Golestanian , Philip Bittihn

The collective motion of epithelial cells is a fundamental biological process which plays a significant role in embryogenesis, wound healing and tumor metastasis. While it has been broadly investigated for over a decade both in vivo and in…

A recently introduced active phase field crystal model describes the formation of ordered resting and traveling crystals in systems of self-propelled particles. Increasing the active drive, a resting crystal can be forced to perform…

Soft Condensed Matter · Physics 2014-02-18 Andreas M. Menzel , Takao Ohta , Hartmut Löwen

Active glassy matter has recently emerged as a novel class of non-equilibrium soft matter, combining energy-driven, active particle movement with dense and disordered glass-like behavior. Here we review the state-of-the-art in this field…

Soft Condensed Matter · Physics 2019-10-02 Liesbeth M. C. Janssen

Flocking, as paradigmatically exemplified by birds, is the coherent collective motion of active agents. As originally conceived, flocking emerges through alignment interactions between the agents. Here, we report that flocking can also…

Soft Condensed Matter · Physics 2024-07-16 Suchismita Das , Matteo Ciarchi , Ziqi Zhou , Jing Yan , Jie Zhang , Ricard Alert

Biological and robotic systems often operate in confined environments where material must be gathered without centralized control. Inspired by the effective collection strategies of aquatic worms (Lumbriculus variegatus and Tubifex…

Soft Condensed Matter · Physics 2025-05-09 R. Sinaasappel , K. R. Prathyusha , H. Tuazon , E. Mirzahossein , P. Illien , S. Bhamla , A. Deblais

The onset of polar flocking in active matter is discontinuous, akin to gas-liquid phase transitions, except that the steady state exhibits microphase separation into polar clusters. While these features have been observed in theoretical…

Soft Condensed Matter · Physics 2021-03-31 Lorenz Huber , Timo Krüger , Erwin Frey

Self-organization is the generation of order out of local interactions in non-equilibrium [1]. It is deeply connected to all fields of science from physics, chemistry to biology where functional living structures self-assemble[2] and…

Soft Condensed Matter · Physics 2018-10-24 Utsab Khadka , Viktor Holubec , Haw Yang , Frank Cichos

We investigate the susceptibility of long-range ordered phases of two-dimensional dry aligning active matter to population disorder, taken in the form of a distribution of intrinsic individual chiralities. Using a combination of…

Soft Condensed Matter · Physics 2021-12-15 Bruno Ventejou , Hugues Chaté , Raul Montagne , Xia-qing Shi

Active matter systems encompass both natural and artificially created systems consisting of numerous active particles. These particles actively consume energy to propel themselves or exert mechanical forces, leading to intricate behaviors…

Statistical Mechanics · Physics 2026-05-12 Mintu Karmakar

We study the spatio-temporal dynamics of a model of polar active fluid in two dimensions. The system exhibits a transition from an isotropic to a polarized state as a function of density. The uniform polarized state is, however, unstable…

Soft Condensed Matter · Physics 2011-12-08 Luca Giomi , M. Cristina Marchetti

The emergence of collective motion, also known as flocking or swarming, in groups of moving individuals who orient themselves using only information from their neighbors is a very general phenomenon that is manifested at multiple spatial…

Statistical Mechanics · Physics 2016-04-26 David A. Quint , Ajay Gopinathan

Emergence of regular spatial patterns is a hallmark in living matter ranging from subcellular organelles to developing embryos and to ecosystems. Mechanisms for the formation of ordered spatial patterns in biology often require chemical…

Biological Physics · Physics 2024-03-15 Haoran Xu , Yilin Wu

Here we show that coupling to curvature has profound effects on collective motion in active systems, leading to patterns not observed in flat space. Biological examples of such active motion in curved environments are numerous: curvature…

Soft Condensed Matter · Physics 2015-03-05 Rastko Sknepnek , Silke Henkes

Recent studies aimed at investigating artificial analogs of bacterial colonies have shown that low-density suspensions of self-propelled particles confined in two dimensions can assemble into finite aggregates that merge and split, but have…

Soft Condensed Matter · Physics 2015-06-17 B. M. Mognetti , A. Šarić , S. Angioletti-Uberti , A. Cacciuto , C. Valeriani , D. Frenkel

One of the most well known features of active matter is the tendencey of self-propelled particles to undergo system-wide collective motion. With low enough rotational noise or high enough global density, these systems spontaneously break…

Soft Condensed Matter · Physics 2024-06-21 Caleb J. Anderson Olivier Dauchot , Alberto Fernandez-Nieves

The self-organised motion of vast numbers of creatures in a single direction is a spectacular example of emergent order. We recreate this phenomenon using actuated non-living components. We report here that millimetre-sized tapered rods,…

Soft Condensed Matter · Physics 2014-09-23 Nitin Kumar , Harsh Soni , Sriram Ramaswamy , A. K. Sood