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Many microswimmers are able to swim through viscous fluids by employing periodic non-reciprocal deformations of their appendages. Here we use a simple microswimmer model inspired by swimming biflagellates which consists of a spherical cell…

Soft Condensed Matter · Physics 2025-08-22 Sridhar Bulusu , Andreas Zöttl

Living cells and tissues can generate complex patterns of electric fields and fluid flows which can play important role in physiology. Both, fields and flows are rooted in ion transport across biological interfaces: cell membranes and…

Soft Condensed Matter · Physics 2026-05-22 Amit Singh Vishen , Ahandeep Manna , Frank Jülicher

Cilia and flagella are actively bending slender organelles, performing functions such as motility, feeding and embryonic symmetry breaking. We review the mechanics of viscous-dominated microscale flow, including time-reversal symmetry, drag…

Quantitative Methods · Quantitative Biology 2013-09-06 Thomas D. Montenegro-Johnson , Andrew A. Smith , David J. Smith , Daniel Loghin , John R. Blake

Microswimmers can acquire information on the surrounding fluid by sensing mechanical queues. They can then navigate in response to these signals. We analyse this navigation by combining deep reinforcement learning with direct numerical…

Fluid Dynamics · Physics 2023-06-21 Krongtum Sankaewtong , John J. Molina , Matthew S. Turner , Ryoichi Yamamoto

An outstanding characteristic of porous media, desired in many applications, is the large surface area, which facilitates solid-fluid interactions, making porous media an extreme case in colloid and interface science. In two-fluid systems,…

Fluid Dynamics · Physics 2025-10-23 Steffen Berg , Ryan T. Armstrong , Maja Rücker , Alex Hansen , Signe Kjelstrup , Dick Bedeaux

Unlike coffee and cream that homogenize when stirred, growing micro-organisms (e.g., bacteria, baker's yeast) can actively kill each other and avoid mixing. How do such antagonistic interactions impact the growth and survival of competing…

Populations and Evolution · Quantitative Biology 2025-04-01 Jonathan Bauermann , Roberto Benzi , David R. Nelson , Suraj Shankar , Federico Toschi

We study how hydrodynamic interactions affect the collective behaviour of active particles suspended in a fluid at high concentrations, with particular attention to lubrication forces which appear when the particles are very close to one…

Soft Condensed Matter · Physics 2018-05-29 Natsuhiko Yoshinaga , Tanniemola B. Liverpool

Collective vertical swimming may generate aggregate-scale flows that contribute to mixing and transport in stratified environments. The strength of these flows depends not only on swimmer behavior but also on environmental properties. Here…

Fluid Dynamics · Physics 2026-05-11 Nina Mohebbi , John O. Dabiri

Bacteria on surfaces exhibit collective behaviors, such as active turbulence and active stresses, which result from their motility, growth, and interactions with their local surroundings. However, interfacial deformations on soft surfaces…

Soft Condensed Matter · Physics 2022-09-13 Vincent Hickl , Gabriel Juarez

Membrane pores are implicated in several critical functions, including cell fusion and the transport of signaling molecules for intercellular communication. However, these structural features are often difficult to probe directly. Droplet…

Soft Condensed Matter · Physics 2025-11-13 Nivedina A. Sarma , David A. King , Xuefei Wu , Brett A. Helms , Paul D. Ashby , Thomas P. Russell , Ahmad K. Omar

We explore a mechanism of the anomalous rheology of active suspensions by hydrodynamic simulations using model pusher swimmers. Our simulations demonstrate that hydrodynamic interactions under shear flow systematically orient swimmers along…

Soft Condensed Matter · Physics 2023-12-19 Haruki Hayano , Akira Furukawa

Biofilms are bacterial aggregates that grow on moist surfaces. Thin homogeneous biofilms naturally formed on the walls of conducts may serve as biosensors, providing information on the status of microsystems (MEMS) without disrupting them.…

Soft Condensed Matter · Physics 2024-01-17 Ana Carpio , Baldvin Einarsson , David R. Espeso

The recent interest in using microorganisms for biofuels is motivation enough to study bioconvection and cell dispersion in tubes subject to imposed flow. To optimize light and nutrient uptake, many microorganisms swim in directions biased…

Soft Condensed Matter · Physics 2010-10-05 O. A. Croze , E. E. Ashraf , M. A. Bees

This study systematically investigates the dynamics of the bacterial transition from free-swimming to surface adhesion, a process characterized by both height $h$ and inclination angle $\Psi$. The surface entrapment process is divided into…

Soft Condensed Matter · Physics 2025-06-25 Baopi Liu , Bowen Jin , Ning An

Cells swimming in confined environments are attracted by surfaces. We measure the steady-state distribution of smooth-swimming bacteria (Escherichia coli) between two glass plates. In agreement with earlier studies, we find a strong…

Soft Condensed Matter · Physics 2008-10-02 Allison P. Berke , Linda Turner , Howard C. Berg , Eric Lauga

Bacteria in biofilms form complex structures and can collectively migrate within mobile aggregates, which is referred to as swarming. This behavior is influenced by a combination of various factors, including morphological characteristics…

Soft Condensed Matter · Physics 2024-01-19 Bohan Wu-Zhang , Dmitry A. Fedosov , Gerhard Gompper

Unicellular microscopic organisms living in aqueous environments outnumber all other creatures on Earth. A large proportion of them are able to self-propel in fluids with a vast diversity of swimming gaits and motility patterns. In this…

Biological Physics · Physics 2021-07-14 Marcos F. Velho Rodrigues , Maciej Lisicki , Eric Lauga

Controlling the swimming behavior of bacteria is crucial, for example, to prevent contamination of ducts and catheters. We show the bacteria modeled by deformable microswimmers can accumulate in flows through straight microchannels either…

Soft Condensed Matter · Physics 2022-03-03 Winfried Schmidt , Igor S. Aranson , Walter Zimmermann

Turbulence is ubiquitous, from oceanic currents to small-scale biological and quantum systems. Self-sustained turbulent motion in microbial suspensions presents an intriguing example of collective dynamical behavior amongst the simplest…

Peritrichous bacteria synchronize and bundle their flagella to actively swim while disruption of the bundle leads to tumbling. It is still not known whether the number of flagella represents an evolutionary adaptation towards optimizing…

Biological Physics · Physics 2020-04-21 Javad Najafi , M. Reza Shaebani , Thomas John , Florian Altegoer , Gert Bange , Christian Wagner