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The physical chemistry of liquid-liquid phase separation (LLPS) of polymer solutions bears directly on the assembly of biologically functional droplet-like bodies from proteins and nucleic acids. These biomolecular condensates include…

Biomolecules · Quantitative Biology 2020-01-31 Yi-Hsuan Lin , Jacob P. Brady , Hue Sun Chan , Kingshuk Ghosh

A theory for sequence dependent liquid-liquid phase separation (LLPS) of intrinsically disordered proteins (IDPs) in the study of biomolecular condensates is formulated by extending the random phase approximation (RPA) and field-theoretic…

Biomolecules · Quantitative Biology 2022-11-28 Jonas Wessén , Suman Das , Tanmoy Pal , Hue Sun Chan

Biomolecular condensates undergirded by phase separations of proteins and nucleic acids serve crucial biological functions. To gain physical insights into their genetic basis, we study how liquid-liquid phase separation (LLPS) of…

Biomolecules · Quantitative Biology 2018-12-03 Suman Das , Alan Amin , Yi-Hsuan Lin , Hue Sun Chan

Endeavoring toward a transferable, predictive coarse-grained explicit-chain model for biomolecular condensates underlain by liquid-liquid phase separation (LLPS), we conducted multiple-chain simulations of the N-terminal intrinsically…

Biomolecules · Quantitative Biology 2020-12-10 Suman Das , Yi-Hsuan Lin , Robert M. Vernon , Julie D. Forman-Kay , Hue Sun Chan

Phase separation of intrinsically disordered proteins is important for the formation of membraneless organelles, or biomolecular condensates, which play key roles in the regulation of biochemical processes within cells. In this work, we…

Soft Condensed Matter · Physics 2020-03-18 Antonia Statt , Helena Casademunt , Clifford P. Brangwynne , Athanassios Z. Panagiotopoulos

The condensation of biomolecules into biomolecular condensates via liquid-liquid phase separation (LLPS) is a ubiquitous mechanism that drives cellular organization. To enable these functions, biomolecules have evolved to drive LLPS and…

Biomolecules · Quantitative Biology 2020-03-04 Erik W. Martin , Jesse B. Hopkins , Tanja Mittag

We investigate liquid-liquid phase separation (LLPS) and interfacial properties of two LLPS modes: associative (ALLPS) and segregative (SLLPS). Analytical expressions for the critical point (CP) and binodal boundaries are derived and show…

Soft Condensed Matter · Physics 2025-10-30 Remco Tuinier , Alvaro Gonzalez Garcia

Liquid liquid phase separation (LLPS) of proteins is an intracellular process that is widely used by cells for many purposes. In living cells (in vivo), LLPS occurs in complex and crowded environments. Amino acids (AAs) are vital components…

Liquid-liquid phase separation (LLPS) of protein solutions is governed by highly complex protein-protein interactions. Nevertheless, it has been suggested that based on the extended law of corresponding states (ELCS), as proposed for…

Soft Condensed Matter · Physics 2023-02-08 Jan Hansen , Stefan U. Egelhaaf , Florian Platten

The precise control of liquid-liquid phase separation (LLPS) is the key to developing cutting-edge technologies that benefit diverse disciplines. Fluid flow was found to be capable of controlling the structure and effective temperature of…

Fluid Dynamics · Physics 2024-07-04 Yulin Li , Tong Zhou , Yanyu Li , Qi Zhang , Zhihong You

The phase behavior is investigated for systems composed of a large number of macromolecular components N, with N greater or equal to 2. Liquid-liquid phase separation is modelled using a virial expansion up to the second order of the…

Soft Condensed Matter · Physics 2024-05-29 Arjen Bot , Erik van der Linden , Paul Venema

Liquid-liquid phase separation is the mechanism underlying the formation of biomolecular condensates. Disordered protein regions often drive phase separation, but molecular interactions of disordered protein regions are not well understood,…

Biomolecules · Quantitative Biology 2020-08-10 Wade Borcherds , Anne Bremer , Madeleine B. Borgia , Tanja Mittag

The energy landscape dictates pathways and outcomes in supramolecular selfassembly, yet harnessing it from the nano to the macro scales remains a major challenge. Here, we demonstrate liquid liquid phase separation (LLPS) as a powerful tool…

The paradigm that the primary amino acid sequence prescribes structure and thus function has for a long time been central to the understanding of protein science. Though the theory is supported by the behaviour of most structured proteins,…

Biological Physics · Physics 2022-12-19 Rickie Xian , Sarah Rauscher

Membraneless droplets or liquid condensates formed via liquid-liquid phase separation (LLPS) play a pivotal role in cell biology and hold potential for biomedical engineering. While membraneless droplets are often studied in the context of…

Soft Condensed Matter · Physics 2025-11-07 Chenxi Liu , Ding Cao , Siyu Liu , Yilin Wu

The assembly of functional biomolecular condensates often involves liquid-liquid phase separation (LLPS) of proteins with multiple modular domains, which can be folded or conformationally disordered to various degrees. To understand the…

Biomolecules · Quantitative Biology 2022-01-11 Yi-Hsuan Lin , Haowei Wu , Bowen Jia , Mingjie Zhang , Hue Sun Chan

Liquid-liquid phase separation is now recognized as a common mechanism for regulating enzyme activity in cells. Insights from studies in cells are complemented by in vitro studies aimed at developing better understanding of mechanisms…

Biomolecules · Quantitative Biology 2020-10-23 Brian G. O'Flynn , Tanja Mittag

The formation of biomolecular condensates inside cells often involve intrinsically disordered proteins (IDPs), and several of these IDPs are also capable of forming droplet-like dense assemblies on their own, through liquid-liquid phase…

Biological Physics · Physics 2020-02-26 Daniel Nilsson , Anders Irbäck

Proteins can form droplets via liquid-liquid phase separation (LLPS) in cells. Recent experiments demonstrate that LLPS is qualitatively different on two-dimensional (2d) surfaces compared to three-dimensional (3d) solutions. In this paper,…

Quantitative Methods · Quantitative Biology 2024-08-29 Jinyoung Kim , Sean D. Lawley , Jinsu Kim

Intrinsically disordered proteins (IDPs) are important for biological functions. In contrast to folded proteins, molecular recognition among certain IDPs is "fuzzy" in that their binding and/or phase separation are stochastically governed…

Biomolecules · Quantitative Biology 2020-08-10 Alan N. Amin , Yi-Hsuan Lin , Suman Das , Hue Sun Chan
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