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In isotropic fluids like water, micrometer-scale swimmers have evolved swim strokes to translate despite their tiny size. As described by Purcell in his Scallop Theorem, reciprocal motions, like those performed by a scallop, cannot drive…

We reconsider fluid dynamics for a self-propulsive swimmer in Stokes flow. With an exact definition of deformation of a swimmer, a proof is given to Purcell's scallop theorem including the body rotation. The breakdown of the theorem due to…

Fluid Dynamics · Physics 2011-08-01 Kenta Ishimoto , Michio Yamada

By synergistically combining modeling, simulation and experiments, we show that there exists a regime of self-propulsion in which the inertia in the fluid dynamics can be separated from that of the swimmer. This is demonstrated by the…

Due to the kinematic reversibility of Stokes flow, a body executing a reciprocal motion (a motion in which the sequence of body configurations remains identical under time reversal) cannot propel itself in a viscous fluid in the limit of…

Soft Condensed Matter · Physics 2009-04-30 David Gonzalez-Rodriguez , Eric Lauga

E. M. Purcell showed that a body has to perform non-reciprocal motion in order to propel itself in a highly viscous environment. The swimmer with one degree of freedom is bound to do reciprocal motion, whereby the center of mass of the…

Soft Condensed Matter · Physics 2017-08-24 Priyanka Choudhary , Subhayan Mandal , Sujin B. Babu

Purcell's scallop theorem defines the type of motions of a solid body - reciprocal motions - which cannot propel the body in a viscous fluid with zero Reynolds number. For example, the flapping of a wing is reciprocal and, as was recently…

Soft Condensed Matter · Physics 2008-10-02 Eric Lauga

In this article, we are interested in studying locomotion strategies for a class of shape-changing bodies swimming in a fluid. This class consists of swimmers subject to a particular linear dynamics, which includes the two most investigated…

Mathematical Physics · Physics 2010-08-09 Alexandre Munnier , Thomas Chambrion

To achieve propulsion at low Reynolds number, a swimmer must deform in a way that is not invariant under time-reversal symmetry; this result is known as the scallop theorem. We show here that there is no many-scallop theorem. We demonstrate…

Soft Condensed Matter · Physics 2008-10-02 Eric Lauga , Denis Bartolo

Locomotion on small scales is dominated by the effects of viscous forces and, as a result, is subject to strong physical and mathematical constraints. Following Purcell's statement of the scallop theorem which delimitates the types of…

Biological Physics · Physics 2011-08-30 Eric Lauga

In a world without inertia, Purcell's scallop theorem states that in a Newtonian fluid a time-reversible motion cannot produce any net force or net flow. Here we consider the extent to which the nonlinear rheological behavior of…

Fluid Dynamics · Physics 2010-04-09 On Shun Pak , Eric Lauga

Purcell's scallop theorem states that swimmers deforming their shapes in a time-reversible manner ("reciprocal" motion) cannot swim. Using numerical simulations and theoretical calculations we show here that in a fluctuating environment,…

Soft Condensed Matter · Physics 2011-08-30 Eric Lauga

We investigate the way in which oscillating dumb-bells, a simple microscopic model of apolar swimmers, move at low Reynold's number. In accordance with Purcell's Scallop Theorem a single dumb-bell cannot swim because its stroke is…

Soft Condensed Matter · Physics 2009-11-13 G. P. Alexander , J. M. Yeomans

In this paper we focus on a two-link swimmer called scallop which moves changing dynamics between two fluids regimes. We address and solve explicitly two optimal control problems, the minimum time one and the minimum quadratic cost needed…

Optimization and Control · Mathematics 2019-03-05 Rosario Maggistro , Marta Zoppello

Swimming and pumping at low Reynolds numbers are subject to the "Scallop theorem", which states that there will be no net fluid flow for time reversible motions. Living organisms such as bacteria and cells are subject to this constraint,…

Soft Condensed Matter · Physics 2009-01-20 M. Leoni , J. Kotar , B. Bassetti , P. Cicuta , M. Cosentino Lagomarsino

Efficient locomotion is important for the evolution of complex life, yet the physical principles selecting specific swimming strokes often remain entangled with biological constraints. In viscous fluids, the scallop theorem constrains the…

Biological Physics · Physics 2026-03-11 Takahiro Kanazawa , Kenta Ishimoto , Kyogo Kawaguchi

This study investigates the dynamics and controllability of a Purcell three-link microswimmer equipped with passive elastic torsional coils at its joints. By controlling the spontaneous curvature, we analyse the swimmers motion using both…

Mathematical Physics · Physics 2025-02-26 Rossella Attanasi , Marta Zoppello , Gaetano Napoli

In Stokes flow, Purcell's scallop theorem forbids objects with time-reversible (reciprocal) swimming strokes from moving. In the presence of inertia, this restriction is eased and reciprocally deforming bodies can swim. A number of recent…

Fluid Dynamics · Physics 2022-11-30 Nicholas J. Derr , Thomas Dombrowski , Chris H. Rycroft , Daphne Klotsa

In this paper we investigate different strategies to overcome the scallop theorem. We will show how to obtain a net motion exploiting the fluid's type change during a periodic deformation. We are interested in two different models: in the…

Dynamical Systems · Mathematics 2016-11-08 Fabio Bagagiolo , Rosario Maggistro , Marta Zoppello

Fluid-based locomotion at low Reynolds number is subject to the constraints of the scallop theorem, which dictate that body kinematics identical under a time-reversal symmetry (in particular, those with a single degree of freedom) cannot…

Biological Physics · Physics 2013-03-13 Gregory L. Wagner , Eric Lauga

Self-propulsion at low Reynolds number is notoriously restricted, a concept that is commonly known as the "scallop theorem". Here we present a truly self-propelled swimmer (force- and torque- free) that, while unable to swim in a Newtonian…

Fluid Dynamics · Physics 2021-11-23 Laurel A. Kroo , Jeremy P. Binagia , Noah Eckman , Manu Prakash , Eric S. G. Shaqfeh
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