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Related papers: Three-Dimensional Loop Extrusion

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Condensation of hundreds of mega-base-pair-long human chromosomes in a small nuclear volume is a spectacular phenomenon. This process is driven by the formation of chromosome loops. ATP consuming motor, condensin, interacts with chromatin…

Soft Condensed Matter · Physics 2021-10-27 Ryota Takaki , Atreya Dey , Guang Shi , Dave Thirumalai

DNA loop extrusion is arguably one of the most important players in genome organization. The precise mechanism by which loop extruding factors (LEFs) work is still unresolved and much debated. One of the major open questions in this field…

Soft Condensed Matter · Physics 2023-09-18 Andrea Bonato , Jae-Won Jang , Kyoung-Wook Moon , Davide Michieletto , Je-Kyung Ryu

The spatial organization of DNA involves DNA loop extrusion and the formation of protein-DNA condensates. While the significance of each process is increasingly recognized, their interplay remains unexplored. Using molecular dynamics…

Biological Physics · Physics 2025-01-31 Ryota Takaki , Yahor Savich , Jan Brugués , Frank Jülicher

Structural Maintenance Complexes (SMC) are energy consuming motors that are important in folding the genome by loop extrusion (LE) in all stages of the cell cycle. Single molecule magnetic tweezer pulling experiments have revealed that…

Soft Condensed Matter · Physics 2026-05-07 Atreya Dey , Guang Shi , Ryota Takaki , D. Thirumalai

Active loop extrusion - the process of formation of dynamically growing chromatin loops due to the motor activity of DNA-binding protein complexes - is firmly established mechanism responsible for chromatin spatial organization at different…

Statistical Mechanics · Physics 2023-01-11 Sergey Belan , Vladimir Parfenyev

Chromatin loop extrusion by Structural Maintenance of Chromosome (SMC) complexes is thought to underlie intermediate-scale chromatin organization inside cells. Motivated by a number of experiments suggesting that nucleosomes may block loop…

Biological Physics · Physics 2021-09-01 Hao Yan , Ivan Surovtsev , Jessica F Williams , Mary Lou P Bailey , Megan C King , Simon G J Mochrie

Chromatin loop extrusion is a popular model for the formation of CTCF loops and topological domains. Recent HiC data have revealed a strong bias in favour of a particular arrangement of the CTCF binding motifs that stabilize loops, and…

Biological Physics · Physics 2020-10-07 C. A. Brackley , J. Johnson , D. Michieletto , A. N. Morozov , M. Nicodemi , P. R. Cook , D. Marenduzzo

An active loop-extrusion mechanism is regarded as the main out--of--equilibrium mechanism responsible for the structuring of megabase-sized domains in chromosomes. We developed a model to study the dynamics of the chromosome fibre by…

Biomolecules · Quantitative Biology 2020-09-30 M. Crippa , Y. Zhan , G. Tiana

Motivated by the recent experimental observations of the DNA loop extrusion by protein motors, in this paper we investigate the statistical properties of the growing polymer loops within the ideal chain model. The loop conformation is…

Statistical Mechanics · Physics 2021-04-30 Dmitry Starkov , Vladimir Parfenyev , Sergey Belan

Transcriptional noise, or heterogeneity, is important in cellular development and in disease. The molecular mechanisms driving it are, however, elusive and ill-understood. Here, we use computer simulations to explore the role of 3D…

Biological Physics · Physics 2025-10-13 Michael Chiang , Cleis Battaglia , Giada Forte , Chris A. Brackley , Nick Gilbert , Davide Marenduzzo

In many biochemical processes, proteins bound to DNA at distant sites are brought into close proximity by loops in the underlying DNA. For example, the function of some gene-regulatory proteins depends on such DNA looping interactions. We…

Quantitative Methods · Quantitative Biology 2009-11-13 John F Beausang , Philip C Nelson

Loop extrusion is one of the main processes shaping chromosome organisation across the cell cycle, yet its role in regulating DNA entanglement and nucleoplasm viscoelasticity remains overlooked. We simulate entangled solutions of linear…

Soft Condensed Matter · Physics 2025-02-03 Filippo Conforto , Yair Augusto Gutierrez Fosado , Davide Michieletto

We propose a model for the formation of chromatin loops based on the diffusive sliding of a DNA-bound factor which can dimerise to form a molecular slip-link. Our slip-links mimic the behaviour of cohesin-like molecules, which, along with…

Biological Physics · Physics 2017-10-04 C. A. Brackley , J. Johnson , D. Michieletto , A. N. Morozov , M. Nicodemi , P. R. Cook , D. Marenduzzo

We use Brownian dynamics simulations to study the formation of chromatin loops through diffusive sliding of slip-link-like proteins, mimicking the behaviour of cohesin molecules. We recently proposed that diffusive sliding is sufficient to…

Soft Condensed Matter · Physics 2020-11-02 Andrea Bonato , Chris A. Brackley , James Johnson , Davide Michieletto , Davide Marenduzzo

Chromosomes are crumpled polymer chains further folded into a sequence of stochastic loops via loop extrusion. While extrusion has been verified experimentally, the particular means by which the extruding complexes bind DNA polymer remains…

Soft Condensed Matter · Physics 2023-06-14 Kirill Polovnikov , Bogdan Slavov

We develop a kinetic theory of cohesin-driven loop extrusion on a disordered chromatin track with transient barriers. In the stationary state, the mean loop size is shown to obey a universal law determined by the bare processivity and a…

Soft Condensed Matter · Physics 2026-04-20 A. Chervinskaya , R. Metzler , K. E. Polovnikov

Autoencoding has achieved great empirical success as a framework for learning generative models for natural images. Autoencoders often use generic deep networks as the encoder or decoder, which are difficult to interpret, and the learned…

Computer Vision and Pattern Recognition · Computer Science 2023-02-21 Xili Dai , Ke Chen , Shengbang Tong , Jingyuan Zhang , Xingjian Gao , Mingyang Li , Druv Pai , Yuexiang Zhai , XIaojun Yuan , Heung-Yeung Shum , Lionel M. Ni , Yi Ma

We calculate the probability of DNA loop formation mediated by regulatory proteins such as Lac repressor (LacI), using a mathematical model of DNA elasticity. Our model is adapted to calculating quantities directly observable in Tethered…

Biomolecules · Quantitative Biology 2009-11-13 Kevin B. Towles , John F. Beausang , Hernan G. Garcia , Rob Phillips , Philip C. Nelson

Chromatin fibers composed of DNA and proteins fold into consecutive loops to form rod-shaped chromosome in mitosis. Although the loop growth dynamics is investigated in several studies, its detailed processes are unclear. Here, we describe…

Soft Condensed Matter · Physics 2022-05-11 Hiroshi Yokota , Masashi Tachikawa

We consider a general class of (intersecting) loop models in D dimensions, including those related to high-temperature expansions of well-known spin models. We find that the loop models exhibit some interesting features - often in the…

Statistical Mechanics · Physics 2007-05-23 L. Chayes , Leonid P. Pryadko , Kirill Shtengel
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