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Many microorganisms find themselves immersed in fluids displaying non-Newtonian rheological properties such as viscoelasticity and shear-thinning viscosity. The effects of viscoelasticity on swimming at low Reynolds numbers have already…

Fluid Dynamics · Physics 2015-11-10 Charu Datt , Lailai Zhu , Gwynn J. Elfring , On Shun Pak

We theoretically investigate self-oscillating waves of an active material, which have recently been introduced as a non-symmetric part of the elastic moduli, termed odd elasticity. Using Purcell's three-link swimmer model, we reveal that an…

Soft Condensed Matter · Physics 2022-06-22 Kenta Ishimoto , Clément Moreau , Kento Yasuda

We explore hydrodynamic interactions between microswimmers and corrugated, or rough, surfaces, as found often in biological systems and microfluidic devices. Using the Lorentz reciprocal theorem for viscous flows we derive exact expressions…

Soft Condensed Matter · Physics 2022-08-10 Christina Kurzthaler , Howard A. Stone

Fish typically swim by periodic bending of their bodies. Bending seems to follow a universal rule; it occurs at about one-third from the posterior end of the fish body with a maximum bending angle of about $30^o$. However, the hydrodynamic…

Fluid Dynamics · Physics 2021-12-22 Haotian Hang , Sina Heydari , John H. Costello , Eva Kanso

Swimming in low-Reynolds-number fluids requires the breaking of time-reversal symmetry and centrosymmetry. Microswimmers, often with asymmetric shapes, exhibit nonreciprocal motions or exploit nonequilibrium processes to propel. The role of…

Soft Condensed Matter · Physics 2023-07-20 Sung-Jo Kim , Žiga Kos , Eujin Um , Joonwoo Jeong

The design of artificial microswimmers is often inspired by the strategies of natural microorganisms. Many of these creatures exploit the fact that elasticity breaks the time-reversal symmetry of motion at low Reynolds numbers, but this…

Experiments are reported on intermittent swimming motions. Water tunnel experiments on a nominally two-dimensional pitching foil show that the mean thrust and power scale linearly with the duty cycle, from a value of 0.2 all the way up to…

Fluid Dynamics · Physics 2018-08-29 Daniel Floryan , Tyler Van Buren , Alexander J. Smits

We uncover subtle and previously unexplored phenomena arising from the interplay of nonlinearity and nonreciprocity in topological mechanical metamaterials. We study a nonreciprocal topological Klein-Gordon chain of asymmetrically coupled…

Disordered Systems and Neural Networks · Physics 2026-02-10 Bertin Many Manda , Vassos Achilleos

Microscale propulsion is integral to numerous biomedical systems, for example biofilm formation and human reproduction, where the surrounding fluids comprise suspensions of polymers. These polymers endow the fluid with non-Newtonian…

Fluid Dynamics · Physics 2018-06-28 David A. Gagnon , Thomas D. Montenegro-Johnson

Low Reynolds number direct simulations of large populations of hydrodynamically interacting swimming particles confined between planar walls are performed. The results of simulations are compared with a theory that describes dilute…

Fluid Dynamics · Physics 2015-05-13 Juan P. Hernandez-Ortiz , Patrick T. Underhill , Michael D. Graham

We investigate the way in which oscillating dumb-bells, a simple microscopic model of apolar swimmers, move at low Reynold's number. In accordance with Purcell's Scallop Theorem a single dumb-bell cannot swim because its stroke is…

Soft Condensed Matter · Physics 2009-11-13 G. P. Alexander , J. M. Yeomans

The current study presents a systematic investigation of the locomotion performance of a swimmer with a wide range of parameter settings. Two-dimensional simulations with the immersed boundary method are employed for the fluid-structure…

Fluid Dynamics · Physics 2023-12-21 Wenkang Wang , Xu Chu

Non-reciprocal interactions are a generic feature of non-equilibrium systems. We define a non-reciprocal generalization of the kinetic Ising model in one spatial dimension. We solve the model exactly using two different approaches for…

Statistical Mechanics · Physics 2025-04-02 Gabriel Artur Weiderpass , Mayur Sharma , Savdeep Sethi

Motility is a fundamental feature of living matter, encompassing single cells and collective behavior. Such living systems are characterized by non-conservativity of energy and a large diversity of spatio-temporal patterns. Thus,…

Soft Condensed Matter · Physics 2023-10-25 Kenta Ishimoto , Clément Moreau , Kento Yasuda

We calculate the hydrodynamic force on a small spherical, unsteady squirmer moving with a time-dependent velocity in a fluid at rest, taking into account convective and unsteady fluid inertia effects in perturbation theory. Our results…

Fluid Dynamics · Physics 2023-12-20 T. Redaelli , F. Candelier , R. Mehaddi , C. Eloy , B. Mehlig

The use of the reciprocal theorem has been shown to be a powerful tool to obtain the swimming velocity of bodies at low Reynolds number. The use of this method for lower-dimensional swimmers, such as cylinders and sheets, is more…

Fluid Dynamics · Physics 2015-11-10 Gwynn J. Elfring

The shape of a cell influences and it is influenced by interactions with its neighbouring partners. Here, we introduce a coarse-grained model of non-reciprocal interactions between single-cell organisms to study emergent morphologies during…

Soft Condensed Matter · Physics 2025-06-17 Maitane Muñoz-Basagoiti , Michael Wassermair , Miguel Amaral , Buzz Baum , Anđela Šarić

The physical mechanism behind the spontaneous formation of metachronal waves in microtubule arrays in a low Reynolds number fluid has been of interest for the past several years, yet is still not well understood. We present a model…

Subcellular Processes · Quantitative Biology 2019-11-06 Stephen E Martin , Matthew E Brunner , Joshua M Deutsch

The hydrodynamics of a flagellated microorganism is investigated when swimming close to a planar free-slip surface by means of numerical solu- tions of the Stokes equations obtained via a Boundary Element Method. Depending on the initial…

Fluid Dynamics · Physics 2017-03-31 Daniela Pimponi , Mauro Chinappi , Paolo Gualtieri , Carlo Massimo Casciola

Two identical particles driven by the same steady force through a viscous fluid may move relative to one another due to hydrodynamic interactions. The presence or absence of this relative translation has a profound effect on the dynamics of…

Soft Condensed Matter · Physics 2021-05-04 Yulia Sokolov , Haim Diamant