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Self-propelled micron-size particles suspended in a fluid, like bacteria or synthetic microswimmers, are strongly non-equilibrium systems where particle motility breaks the microscopic detailed balance, often resulting in large-scale…

Soft Condensed Matter · Physics 2024-03-15 Dóra Bárdfalvy , Viktor Škultéty , Cesare Nardini , Alexander Morozov , Joakim Stenhammar

A collection of microswimmers immersed in an incompressible fluid is characterised by strong interactions due to the long-range nature of the hydrodynamic fields generated by individual organisms. As a result, suspensions of rear-actuated…

Soft Condensed Matter · Physics 2023-12-19 Viktor Škultéty , Dóra Bárdfalvy , Joakim Stenhammar , Cesare Nardini , Alexander Morozov

Suspensions of rear- and front-actuated microswimmers immersed in a fluid, known respectively as ``pushers'' and ``pullers'', display qualitatively different collective behaviours: beyond a characteristic density, pusher suspensions exhibit…

Soft Condensed Matter · Physics 2020-07-08 Dóra Bárdfalvy , Shan Anjum , Cesare Nardini , Alexander Morozov , Joakim Stenhammar

Microswimmers typically operate in complex environments. In biological systems, often diverse species are simultaneously present and interact with each other. Here, we derive a (time-dependent) particle-scale statistical description, namely…

Soft Condensed Matter · Physics 2019-08-13 Christian Hoell , Hartmut Löwen , Andreas M. Menzel

Active matter exhibits various forms of non-equilibrium states in the absence of external forcing, including macroscopic steady-state currents. Such states are often too complex to be modelled from first principles and our understanding of…

Soft Condensed Matter · Physics 2020-09-17 Viktor Škultéty , Cesare Nardini , Joakim Stenhammar , Davide Marenduzzo , Alexander Morozov

Microorganisms can preferentially orient and move along gradients of a chemo-attractant (i.e., chemotax) while colonies of many microorganisms can collectively undergo complex dynamics in response to chemo-attractants that they themselves…

Biological Physics · Physics 2013-10-30 Enkeleida Lushi , Raymond E. Goldstein , Michael J. Shelley

Suspensions of swimming micro-organisms are known to undergo intricate collective dynamics as a result of hydrodynamic and collision interactions. Micro-swimmers, such as bacteria and micro-algae, naturally live and have evolved in complex…

Fluid Dynamics · Physics 2025-10-31 Yasser Almoteri , Enkeleida Lushi

Microswimmers are active particles of microscopic size that self-propel by setting the surrounding fluid into motion. According to the kind of far-field fluid flow that they induce, they are classified into pushers and pullers. Many studies…

Soft Condensed Matter · Physics 2019-08-01 Giorgio Pessot , Hartmut Löwen , Andreas M. Menzel

Collective behaviour in suspensions of microswimmers is often dominated by the impact of long-ranged hydrodynamic interactions. These phenomena include active turbulence, where suspensions of pusher bacteria at sufficient densities exhibit…

Soft Condensed Matter · Physics 2019-08-27 Dora Bardfalvy , Henrik Nordanger , Cesare Nardini , Alexander Morozov , Joakim Stenhammar

Microorganisms ofter move in confined, disordered environments, where hydrodynamic couplings can modify their transport behavior. Using extensive finite-element simulations, we investigate the dynamics of microswimmers -- modeled as…

Soft Condensed Matter · Physics 2026-03-24 Mirko Residori , Sebastian Aland , Christina Kurzthaler

Self-propelled particles with hydrodynamic interactions (microswimmers) have previously been shown to produce long-range ordering phenomena. Many theoretical explanations for these collective phenomena are connected to instabilities in the…

Soft Condensed Matter · Physics 2017-05-03 Yuzhou Qian , Peter R. Kramer , Patrick T. Underhill

We investigate theoretically the collective dynamics of a suspension of low Reynolds number swimmers that are confined to two dimensions by a thin fluid film. Our model swimmer is characterized by internal degrees of freedom which locally…

Biological Physics · Physics 2015-05-19 M. Leoni , T. B. Liverpool

Systems of motile microorganisms exhibit a multitude of collective phenomena, including motility-induced phase separation and turbulence. Sensing of the environment and adaptation of movement plays an essential role in the emergent…

Soft Condensed Matter · Physics 2024-06-26 Segun Goh , Elmar Westphal , Roland G. Winkler , Gerhard Gompper

Suspensions of unicellular microswimmers such as flagellated bacteria or motile algae exhibit spontaneous density heterogeneities at large enough concentrations. Based on the relative location of the biological actuation appendages i.e.…

Soft Condensed Matter · Physics 2017-11-27 Fabian Jan Schwarzendahl , Marco G. Mazza

Hydrodynamical interactions of active micro-particles are pervasive in our planet's fluid environments. Hence, understanding the interactions of these self-propelled particles is essential for science and engineering. In this paper the…

Soft Condensed Matter · Physics 2022-12-28 Yonatan Ashenafi

The complex formations exhibited by schooling fish have long been the object of fascination for biologists and physicists. However, the physical and sensory mechanisms leading to organized collective behavior remain elusive. On the physical…

Fluid Dynamics · Physics 2025-07-09 Anand U. Oza , Eva Kanso , Michael J. Shelley

In this review we discuss the recent progress in the simulation of soft active matter systems and in particular the hydrodynamics of microswimmers using the method of multiparticle collision dynamics, which solves the hydrodynamic flows…

Soft Condensed Matter · Physics 2020-09-03 Andreas Zöttl

Active matter has been the object of huge amount of research in recent years for its important fundamental and applicative properties. In this paper we investigate active suspensions of micro-swimmers through direct numerical simulation, so…

Soft Condensed Matter · Physics 2021-02-03 Sergio Chibbaro , Astrid Decoene , Sebastien Martin , Fabien Vergnet

Microswimmers often exhibit surprising patterns due to the nonequilibrium nature of their dynamics. Collectively, suspensions of microswimmers appear as a liquid whose properties set it apart from its passive counterpart. To understand the…

Soft Condensed Matter · Physics 2019-05-22 Fabian Jan Schwarzendahl , Marco G. Mazza

We explore the mechanism underlying the anomalous rheology of puller-type microswimmer suspensions through direct hydrodynamic simulations. Puller-type swimmers generate contractile flow fields along their swimming direction, leading to…

Soft Condensed Matter · Physics 2025-03-04 Haruki Hayano , Akira Furukawa
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