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Living systems operate far from thermodynamic equilibrium. Enzymatic activity can induce broken detailed balance at the molecular scale. This molecular scale breaking of detailed balance is crucial to achieve biological functions such as…

Biological Physics · Physics 2018-05-02 F. Gnesotto , F. Mura , J. Gladrow , C. P. Broedersz

It is well known that the kinetics of an intracellular biochemical network is stochastic. This is due to intrinsic noise arising from the random timing of biochemical reactions in the network as well as due to extrinsic noise stemming from…

Subcellular Processes · Quantitative Biology 2019-06-05 Emma M. Keizer , Bjorn Bastian , Robert W. Smith , Ramon Grima , Christian Fleck

When cells measure concentrations of chemical signals, they may average multiple measurements over time in order to reduce noise in their measurements. However, when cells are in a environment that changes over time, past measurements may…

Cell Behavior · Quantitative Biology 2025-03-05 Aparajita Kashyap , Wei Wang , Brian A. Camley

Cells are constantly exposed to fluctuating environmental conditions. External signals are sensed, processed and integrated by cellular signal transduction networks, which translate input signals into specific cellular responses by means of…

Molecular Networks · Quantitative Biology 2012-05-29 Pau Rué , Núria Domedel-Puig , Jordi Garcia-Ojalvo , Antonio J. Pons

Active nematics are materials composed of mobile, elongated particles that can transform energy from the environment into a mechanical motion. Current experimental realizations of the active nematics are of biological origin and include…

Soft Condensed Matter · Physics 2024-06-05 Andrey Sokolov , Jaideep Katuri , Juan J. de Pablo , Alexey Snezhko

Nature presents multiple intriguing examples of processes which proceed at high precision and regularity. This remarkable stability is frequently counter to modelers' experience with the inherent stochasticity of chemical reactions in the…

Molecular Networks · Quantitative Biology 2015-12-31 Andreas Milias-Argeitis , Stefan Engblom , Pavol Bauer , Mustafa Khammash

Mechanochemically active enzymes change their shapes within every turnover cycle. Therefore, they induce circulating flows in the solvent around them and behave as oscillating hydrodynamic force dipoles. Because of non-equilibrium…

Soft Condensed Matter · Physics 2021-12-28 Yuto Hosaka , Shigeyuki Komura , Alexander S. Mikhailov

Even in the steady-state, the number of biomolecules in living cells fluctuates dynamically; and the frequency spectrum of this chemical fluctuation carries valuable information about the mechanism and the dynamics of the intracellular…

Biological Physics · Physics 2018-08-21 Sanggeun Song , Gil-Suk Yang , Seong Jun Park , Ji-Hyun Kim , Jaeyoung Sung

The glassy cytoplasm, crowded with bio-macromolecules, is fluidized in living cells by mechanical energy derived from metabolism. Characterizing the living cytoplasm as a non-equilibrium system is crucial in elucidating the intricate…

Biological Physics · Physics 2023-07-28 K. Umeda , K. Nishizawa , W. Nagao , S. Inokuchi , Y. Sugino , H. Ebata , D. Mizuno

Gene regulatory circuits must contend with intrinsic noise that arises due to finite numbers of proteins. While some circuits act to reduce this noise, others appear to exploit it. A striking example is the competence circuit in Bacillus…

Molecular Networks · Quantitative Biology 2016-04-27 Andrew Mugler , Mark Kittisopikul , Luke Hayden , Jintao Liu , Chris H. Wiggins , Gurol M. Suel , Aleksandra M. Walczak

Phenotypic noise underpins homeostasis and fitness of individual cells. Yet, the extent to which noise shapes cell-to-population properties in microbial active matter remains poorly understood. By quantifying variability in confluent…

Biological Physics · Physics 2023-11-07 Jayabrata Dhar , Anh L. P. Thai , Arkajyoti Ghoshal , Luca Giomi , Anupam Sengupta

One of the most celebrated successes in computational biology is the Hodgkin-Huxley framework for modeling electrically active cells. This framework, expressed through a set of differential equations, synthesizes the impact of ionic…

Neurons and Cognition · Quantitative Biology 2015-05-28 Joshua H. Goldwyn , Eric Shea-Brown

Tissues of living cells are a prime example of active fluids. There is experimental evidence that tissues generate extensile active stress even though their constituting cells are contractile. Fluctuating forces that could result from…

Soft Condensed Matter · Physics 2025-07-01 Mathieu Dedenon , Karsten Kruse

Kinetic-transport equations that take into account the intra-cellular pathways are now considered as the correct description of bacterial chemotaxis by run and tumble. Recent mathematical studies have shown their interest and their…

Analysis of PDEs · Mathematics 2018-06-13 Benoît Perthame , Weiran Sun , Min Tang

In response to a concentration gradient of nutrient, E. coli bacterium modulates the rotational bias of flagellar motors which control its run-and-tumble motion, to migrate towards regions of high nutrient concentration. Presence of…

Cell Behavior · Quantitative Biology 2018-04-04 Subrata Dev , Sakuntala Chatterjee

Single molecular motor kinesin harnesses thermal and non-thermal fluctuations to transport various cargoes along microtubules, converting chemical energy to directed movements. To describe the non-thermal fluctuations generated by the…

Soft Condensed Matter · Physics 2022-12-20 Mengkai Feng , Zhonghuai Hou

The phenomena of stochasticity in biochemical processes have been intriguing life scientists for the past few decades. We now know that living cells take advantage of stochasticity in some cases and counteract stochastic effects in others.…

Quantitative Methods · Quantitative Biology 2011-06-09 Michal Komorowski , Jacek Miekisz , Michael P. H. Stumpf

Eukaryotic cell motility is crucial during development, wound healing, the immune response, and cancer metastasis. Some eukaryotic cells can swim, but cells more commonly adhere to and crawl along the extracellular matrix. We study the…

Cell Behavior · Quantitative Biology 2019-09-02 Melissa H. Mai , Brian A. Camley

Biomembranes wrapping cells and organelles are not only the partitions that separate the insides but also dynamic fields for biological functions accompanied by membrane shape changes. In this review, we discuss the spatiotemporal patterns…

Soft Condensed Matter · Physics 2025-01-22 Hiroshi Noguchi

We used various analytical and numerical techniques to elucidate signal propagation in a small enzymatic cascade which is subjected to external and internal noise. The nonlinear character of catalytic reactions, which underlie protein…

Molecular Networks · Quantitative Biology 2009-11-13 Yueheng Lan , Garegin A. Papoian