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The topological properties of many materials are central to their behavior, with the dynamics of topological defects being particularly important to intrinsically out-of-equilibrium, active materials. In this paper, local manipulation of…

Soft Condensed Matter · Physics 2019-11-01 David P. Rivas , Tyler N. Shendruk , Robert R. Henry , Daniel H. Reich , Robert L. Leheny

We report on the emergence of stable self-propelled bound defects in monolayers of active nematics, which form virtual full-integer topological defects in the form of vortices and asters. Through numerical simulations and analytical…

Soft Condensed Matter · Physics 2020-10-26 Kristian Thijssen , Amin Doostmohammadi

Using novel micro-printing techniques, we develop a versatile experimental setup that allows us to study how lateral confinement tames the active flows and defect properties of the microtubule/kinesin active nematic system. We demonstrate…

Active nematics in two dimensions stir themselves efficiently through internally generated chaotic flows, largely driven by motile $+1/2$ disclinations. We investigate how this tendency toward chaotic fluid stirring can, counterintuitively,…

The structure and dynamics of important biological quasi-two-dimensional systems, ranging from cytoskeletal gels to tissues, are controlled by nematic order, flow, defects and activity. Continuum hydrodynamic descriptions combined with…

Biological Physics · Physics 2024-12-03 Waleed Mirza , Alejandro Torres-Sánchez , Guillermo Vilanova , Marino Arroyo

Topological defects are one of the most conspicuous features of liquid crystals. In two dimensional nematics, they have been shown to behave effectively as particles with both, charge and orientation, which dictate their interactions. Here,…

Soft Condensed Matter · Physics 2021-07-14 Daniel J. G. Pearce , Karsten Kruse

Increasing evidence suggests that active matter exhibits instances of mixed symmetry that cannot be fully described by either polar or nematic formalism. Here, we introduce a minimal model that integrates self-propulsion into the active…

Soft Condensed Matter · Physics 2025-09-03 Niels de Graaf Sousa , Simon Guldager Andersen , Aleksandra Ardaševa , Amin Doostmohammadi

Recent studies of the phase diagram for spherical, purely repulsive, active particles established the existence of a transition from a liquid-like to a solid-like phase analogous to the one observed in colloidal systems at thermal…

Statistical Mechanics · Physics 2020-04-30 Lorenzo Caprini , Claudio Maggi , Umberto Marini Bettolo Marconi , Matteo Paoluzzi , Andrea Puglisi

Collective cell migration governs a range of physiological and pathological processes, from tissue morphogenesis to cancer invasion, in which topological defects arise as an inevitable consequence of frequent cellular rearrangement and…

Soft Condensed Matter · Physics 2026-04-01 Jiusi Zhang , Chung Wing Chan , Bo Li , Rui Zhang

I put forward a continuum theory for active nematic gels, defined as fluids or suspensions of orientable rodlike objects endowed with active dynamics, that is based on symmetry arguments and compatibility with thermodynamics. The starting…

Soft Condensed Matter · Physics 2017-11-15 Stefano S Turzi

Spatiotemporal patterns in multicellular systems are important to understanding tissue dynamics, for instance, during embryonic development and disease. Here, we use a multiphase field model to study numerically the behavior of a…

Soft Condensed Matter · Physics 2025-02-13 Michael Chiang , Austin Hopkins , Benjamin Loewe , M. Cristina Marchetti , Davide Marenduzzo

Colloids dispersed in nematic liquid crystals form topological composites in which colloid-associated defects mediate interactions while adhering to fundamental topological constraints. Better realising the promise of such materials…

Soft Condensed Matter · Physics 2024-04-16 Louise C. Head , Yair A. G. Fosado , Davide Marenduzzo , Tyler N. Shendruk

We show that "dry" active nematics, e.g. collections of shaken elongated granular particles, exhibit large-scale spatiotemporal chaos made of interacting dense, ordered, band-like structures in a parameter region including the linear onset…

Soft Condensed Matter · Physics 2014-09-16 Sandrine Ngo , Anton Peshkov , Igor S. Aranson , Eric Bertin , Francesco Ginelli , Hugues Chaté

Spatially localised defect structures emerge spontaneously in a hydrodynamic description of an active polar fluid comprising polar 'actin' filaments and 'myosin' motor proteins that (un)bind to filaments and exert active contractile…

Biological Physics · Physics 2017-02-22 Kabir Husain , Madan Rao

We establish how active stress globally affects the morphology of disclination lines of a three dimensional active nematic liquid crystal under chaotic flow. Thanks to a defect detection algorithm based on the local nematic orientation, we…

Soft Condensed Matter · Physics 2025-06-30 Pasquale Digregorio , Cecilia Rorai , Ignacio Pagonabarraga , Federico Toschi

Active nematics are dense systems of rodlike particles that consume energy to drive motion at the level of the individual particles. They exist in natural systems like biological tissues and artificial materials such as suspensions of…

Soft Condensed Matter · Physics 2023-12-14 Yunrui Li , Zahra Zarei , Phu N. Tran , Yifei Wang , Aparna Baskaran , Seth Fraden , Michael F. Hagan , Pengyu Hong

Morphogenesis emerges from dynamic feedback among geometry, mechanics, and chemistry; however, disentangling these contributions in living systems remains challenging. Here, we focus on the interplay between geometry and mechanics by…

The growing interest in active nematics and the emerging evidence of the relevance of topological defects in biology asks for reliable data analysis tools to identify, classify and track such defects in simulation and microscopy data. We…

Soft Condensed Matter · Physics 2020-02-10 Dennis Wenzel , Michael Nestler , Sebastian Reuther , Maximilian Simon , Axel Voigt

Defects are a ubiquitous feature of ordered media. They have certain universal features, independent of the underlying physical system, reflecting their topological origins. While the topological properties of defects are robust, they…

Soft Condensed Matter · Physics 2021-02-24 Chiqun Zhang , Amit Acharya , Alan C. Newell , Shankar C. Venkataramani

One of the defining features of active nematics is that above a critical activity the quiescent state becomes unstable to a distorted, flowing one. We show that spatial variations in activity can fundamentally change the nature of this…

Soft Condensed Matter · Physics 2025-12-15 Alexander J. H. Houston , Michael Grinfeld , Geoff McKay , Nigel J. Mottram