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Colonies of the green alga $Volvox$ are spheres that swim through the beating of pairs of flagella on their surface somatic cells. The somatic cells themselves are mounted rigidly in a polymeric extracellular matrix, fixing the orientation…

Soft Condensed Matter · Physics 2016-06-29 T. J. Pedley , D. R. Brumley , R. E. Goldstein

Hydrodynamic synchronization provides a general mechanism for the spontaneous emergence of coherent beating states in independently driven mesoscopic oscillators. A complete physical picture of those phenomena is of definite importance to…

Soft Condensed Matter · Physics 2014-01-29 Nick Koumakis , Roberto Di Leonardo

Recent experiments have shown that the nematode {\it T. aceti} can assemble into collectively undulating groups at the edge of fluid drops. This coordinated state consists of metachronal waves and drives fluid circulation inside the drop.…

Hydrodynamic interaction strongly influences the collective behavior of the microswimmers. With this work, we study the behavior of two hydrodynamically interacting self-propelled chiral swimmers in the low Reynolds number regime,…

Soft Condensed Matter · Physics 2022-11-28 Ruma Maity , P. S. Burada

A system of ferromagnetic particles trapped at a liquid-liquid interface and subjected to a set of magnetic fields (magnetocapillary swimmers) is studied numerically using a hybrid method combining the pseudopotential lattice Boltzmann…

Biological systems are robust to perturbations at both the genetic and environmental levels. Yet, these same perturbations can elicit variation in behavior. The interplay between functional robustness and behavioral variability is…

Biological Physics · Physics 2021-06-03 Veikko Geyer , Jonathon Howard , Pablo Sartori

We investigate the effects of helical swimmer shape (i.e., helical pitch angle and tail thickness) on swimming dynamics in a constant viscosity viscoelastic (Boger) fluid via a combination of particle tracking velocimetry, particle image…

Soft Condensed Matter · Physics 2022-06-01 S. Wu , T. Solano , K. Shoele , H. Mohammadigoushki

When tiny soft ferromagnetic particles are placed along a liquid interface and exposed to a vertical magnetic field, the balance between capillary attraction and magnetic repulsion leads to self-organization into well-defined patterns.…

Soft Condensed Matter · Physics 2020-11-13 Ylona Collard , Galien Grosjean , Nicolas Vandewalle

The motion of a rotating helical body in a viscoelastic fluid is considered. In the case of force-free swimming, the introduction of viscoelasticity can either enhance or retard the swimming speed and locomotive efficiency, depending on the…

Biological Physics · Physics 2013-08-15 Saverio E. Spagnolie , Bin Liu , Thomas R. Powers

Microswimmers are exposed in nature to crowded environments and their transport properties depend in a subtle way on the interaction with obstacles. Here, we investigate a model for a single ideal circle swimmer exploring a two-dimensional…

Biological Physics · Physics 2020-03-17 Oleksandr Chepizhko , Thomas Franosch

Microswimmers display an intriguing ability to navigate through fluids with spatially varying viscosity, a behavior known as viscotaxis, which plays a crucial role in guiding their motion. In this study, we reveal that the orientation…

Fluid Dynamics · Physics 2025-06-25 Takuya Kobayashi , Ryoichi Yamamoto

We study the trajectories of a model microorganism inside three-dimensional channels with square and rectangular cross-sections. Using (i) numerical simulations based on lattice-Boltzmann method, and (ii) analytical expressions using…

Soft Condensed Matter · Physics 2024-08-16 Byjesh N. Radhakrishnan , Ahana Purushothaman , Ranabir Dey , Sumesh P Thampi

In a fluid environment, flagellated microswimmers propel themselves by rotating their flagella. The morphology of these flagella significantly influences forward speed, swimming efficiency, and directional stability, which are critical for…

Fluid Dynamics · Physics 2025-06-25 Baopi Liu , Lu Chen , Wenjun Xu

We study the activity of "living" droplets, which confine 1-6 mesoswimmers in 3D using a superhydrophobic substrate. The swimmers induce oscillations of the droplets at their inherent resonant frequencies, regardless of swimmer size and…

Soft Condensed Matter · Physics 2026-01-30 L. Malik , N. Sharadhi , M. Lamminmäki , R. A. Lara , V. Jokinen , M. Backholm

Microswimmers (planktonic microorganisms or artificial active particles) immersed in a fluid interact with the ambient flow, altering their trajectories. In surface gravity waves, a common goal for microswimmers is vertical migration (e.g.,…

Soft Condensed Matter · Physics 2022-03-01 Kunlin Ma , Nimish Pujara , Jean-Luc Thiffeault

Using multiparticle collision dynamics simulations, we investigate the swimming dynamics, orientational behavior, and hydrodynamic interactions of a model swimmer designed to mimic the isolated flagellar apparatus ($FA$) of Chlamydomonas…

Biological Physics · Physics 2025-07-01 Sai Venkata Ramana Ambadipudi , Albert Bae , Azam Gholami

There is increasing evidence that mammalian cells not only crawl on substrates but can also swim in fluids. To elucidate the mechanisms of the onset of motility of cells in suspension, a model which couples actin and myosin kinetics to…

Biological Physics · Physics 2019-09-18 Alexander Farutin , Jocelyn Etienne , Chaouqi Misbah , Pierre Recho

At high concentration, free swimming nematodes known as vinegar eels ({\it Turbatrix aceti}), collectively exhibit metachronal waves near a boundary. We find that the frequency of the collective traveling wave is lower than that of the…

Soft Condensed Matter · Physics 2023-02-07 A. C. Quillen , A. Peshkov , Esteban Wright , Sonia McGaffigan

Swimming micro-organisms such as bacteria or spermatozoa are typically found in dense suspensions, and exhibit collective modes of locomotion qualitatively different from that displayed by isolated cells. In the dilute limit where…

Biological Physics · Physics 2010-05-02 Sebastien Michelin , Eric Lauga

We employ three numerical methods to explore the motion of low Reynolds number swimmers, modeling the hydrodynamic interactions by means of the Oseen tensor approximation, lattice Boltzmann simulations and multiparticle collision dynamics.…

Soft Condensed Matter · Physics 2007-05-23 David J. Earl , C. M. Pooley , J. F. Ryder , Irene Bredberg , J. M. Yeomans