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External gradients can strongly influence the collective behavior of microswimmers. In this paper, we study the behavior of two hydrodynamically interacting self-propelled chiral swimmers, in the low-Reynolds number regime, under the…

Biological Physics · Physics 2023-01-04 Ruma Maity , P. S. Burada

Biological microswimmers often encounter deformable boundaries in physiological conditions; for instance, the viscoelastic walls of reproductive tract during migration of spermatozoa, or host tissue during early bacterial biofilm formation.…

Soft Condensed Matter · Physics 2025-08-07 Smita S. Sontakke , Aneesha Kajampady , Mohd Suhail Rizvi , Ranabir Dey

We study different types of microswimmers moving in channels with varying cross section and thereby interacting hydrodynamically with the channel walls. Starting from the Smoluchowski equation for a dilute suspension, for which interactions…

Soft Condensed Matter · Physics 2017-05-24 Paolo Malgaretti , Holger Stark

The design of artificial microswimmers is often inspired by the strategies of natural microorganisms. Many of these creatures exploit the fact that elasticity breaks the time-reversal symmetry of motion at low Reynolds numbers, but this…

Microswimmers are encountered in a wide variety of biophysical settings. When interacting with flow fields, they show interesting dynamical features such as trapping, clustering, and preferential orientation. One important step towards the…

Fluid Dynamics · Physics 2020-10-02 José-Agustín Arguedas-Leiva , Michael Wilczek

Microswimmers often use chirality to generate translational movement from rotation motion, exhibiting distinct behaviors in complex fluids compared to simple Newtonian fluids. However, the underlying mechanism remains incompletely…

Soft Condensed Matter · Physics 2024-11-08 Takuya Kobayashi , John J. Molina , Ryoichi Yamamoto

Previous particle-based computer simulations have revealed a significantly more pronounced tendency of spontaneous global polar ordering in puller (contractile) microswimmer suspensions than in pusher (extensile) suspensions. We here…

Soft Condensed Matter · Physics 2018-10-23 Christian Hoell , Hartmut Löwen , Andreas M. Menzel

After colliding with a surface, microswimmers reside there during the detention time. They accumulate and may form complex structures such as biofilms. We introduce a general framework to calculate the distribution of detention times using…

Soft Condensed Matter · Physics 2015-07-21 Konstantin Schaar , Andreas Zöttl , Holger Stark

We study the dynamics of gyrotactic microswimmers suspended in homogeneous and isotropic turbulence by using direct numerical simulations (DNS). The swimmers are characterized by three non-dimensional parameters: their aspect ratio…

Soft Condensed Matter · Physics 2026-04-24 Suraj Kumar Nayak , Vishwanath Shukla , Akshay Bhatnagar

Microswimmers exhibit an intriguing, highly-dynamic collective motion with large-scale swirling and streaming patterns, denoted as active turbulence -- reminiscent of classical high-Reynolds-number hydrodynamic turbulence. Various…

Soft Condensed Matter · Physics 2024-06-03 Kai Qi , Elmar Westphal , Gerhard Gompper , Roland G. Winkler

The net steady state flow pattern of a distorting sphere is studied in the framework of the bilinear theory of swimming at low Reynolds number. It is argued that the starting point of a theory of interacting active particles should be based…

Fluid Dynamics · Physics 2015-05-20 B. U. Felderhof

Swimming, i.e., being able to advance in the absence of external forces by performing cyclic shape changes, is particularly demanding at low Reynolds numbers which is the regime of interest for micro-organisms and micro-robots. We focus on…

Optimization and Control · Mathematics 2012-10-04 François Alouges , Laetitia Giraldi

We present an analytical calculation of the hydrodynamic interaction between two spherical particles near an elastic interface such as a cell membrane. The theory predicts the frequency dependent self- and pair-mobilities accounting for the…

Fluid Dynamics · Physics 2016-08-03 Abdallah Daddi-Moussa-Ider , Stephan Gekle

Most classical work on the hydrodynamics of low-Reynolds-number swimming addresses deterministic locomotion in quiescent environments. Thermal fluctuations in fluids are known to lead to a Brownian loss of the swimming direction. As most…

Fluid Dynamics · Physics 2014-06-18 Mario Sandoval , Navaneeth K. M. , Ganesh Subramanian , Eric Lauga

Self-propelled phoretic swimmers are generally studied in the laminar flow regime, where their low speed renders inertial effects negligible and trajectories highly predictable. This research tackles the challenge of propulsion in the…

Fluid Dynamics · Physics 2026-01-19 Alessandro Foradori , Paolo Bettotti

Recent research has shown that motile cells can adapt their mode of propulsion depending on the environment in which they find themselves. One mode is swimming by blebbing or other shape changes, and in this paper we analyze a class of…

Fluid Dynamics · Physics 2016-10-10 Qixuan Wang , Hans G. Othmer

A dilute non-interacting suspension of micro-swimmers exhibits a finite velocity variance and short-ranged correlations that decay over a swimmer length. For a suspension of interacting straight swimmers, however, pair-interactions leads to…

Soft Condensed Matter · Physics 2020-12-02 Sankalp Nambiar , Piyush Garg , Ganesh Subramanian

We discuss the dynamics of a generalized three-sphere microswimmer in which the spheres are connected by two elastic springs. The natural length of each spring is assumed to undergo a prescribed cyclic change. We analytically obtain the…

Soft Condensed Matter · Physics 2017-08-10 Kento Yasuda , Yuto Hosaka , Mizuki Kuroda , Ryuichi Okamoto , Shigeyuki Komura

The swimming velocity and rate of dissipation of a linear chain consisting of two or three little spheres and a big sphere is studied on the basis of low Reynolds number hydrodynamics. The big sphere is treated as a passive cargo, driven by…

Fluid Dynamics · Physics 2015-06-22 B. U. Felderhof

We present a hydrodynamic study of a monolayer of squirmer model microswimmers confined to a boundary by strong gravity using the simulation method of multi-particle collision dynamics. The squirmers interact with each other via their…

Soft Condensed Matter · Physics 2019-07-23 Jan-Timm Kuhr , Felix Rühle , Holger Stark