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We study how hydrodynamic interactions affect the collective behaviour of active particles suspended in a fluid at high concentrations, with particular attention to lubrication forces which appear when the particles are very close to one…

Soft Condensed Matter · Physics 2018-05-29 Natsuhiko Yoshinaga , Tanniemola B. Liverpool

Few simulations exist for microswimmers near deformable interfaces. Here, we present numerical simulations of the hydrodynamic flows associated with a single microswimmer embedded in a binary fluid mixture. The two fluids demix, separated…

Soft Condensed Matter · Physics 2022-05-24 Chao Feng , John J. Molina , Matthew S. Turner , Ryoichi Yamamoto

The journey of mammalian spermatozoa in nature is well-known to be reliant on their individual motility. Often swimming in crowded microenvironments, the progress of any single swimmer is likely dependent on their interactions with other…

Fluid Dynamics · Physics 2019-09-11 Benjamin J. Walker , Kenta Ishimoto , Eamonn A. Gaffney

We study the trajectories of a model microorganism inside three-dimensional channels with square and rectangular cross-sections. Using (i) numerical simulations based on lattice-Boltzmann method, and (ii) analytical expressions using…

Soft Condensed Matter · Physics 2024-08-16 Byjesh N. Radhakrishnan , Ahana Purushothaman , Ranabir Dey , Sumesh P Thampi

In the study of microscopic flows, self-propulsion has been particularly topical in recent years, with the rise of miniature artificial swimmers as a new tool for flow control, low Reynolds number mixing, micromanipulation or even drug…

Geometric confinements are frequently encountered in soft matter systems and in particular significantly alter the dynamics of swimming microorganisms in viscous media. Surface-related effects on the motility of microswimmers can lead to…

Self-propelled particles can exhibit surprising non-equilibrium behaviors, and how they interact with obstacles or boundaries remains an important open problem. Here we show that chemically propelled micro-rods can be captured, with little…

Soft Condensed Matter · Physics 2014-02-21 Daisuke Takagi , Jeremie Palacci , Adam B. Braunschweig , Michael J. Shelley , Jun Zhang

Microswimmers typically operate in complex environments. In biological systems, often diverse species are simultaneously present and interact with each other. Here, we derive a (time-dependent) particle-scale statistical description, namely…

Soft Condensed Matter · Physics 2019-08-13 Christian Hoell , Hartmut Löwen , Andreas M. Menzel

We study theoretically the behavior of a class of hydrodynamic dipoles. This study is motivated by recent experiments on synthetic and biological swimmers in microfluidic \textit{Hele-Shaw} type geometries. Under such confinement, a…

Fluid Dynamics · Physics 2015-09-30 Eva Kanso , Alan Tsang

Self-propelled micron-size particles suspended in a fluid, like bacteria or synthetic microswimmers, are strongly non-equilibrium systems where particle motility breaks the microscopic detailed balance, often resulting in large-scale…

Soft Condensed Matter · Physics 2024-03-15 Dóra Bárdfalvy , Viktor Škultéty , Cesare Nardini , Alexander Morozov , Joakim Stenhammar

The spatiotemporal dynamics in systems of active self-propelled particles is controlled by the propulsion mechanism in combination with various direct interactions, such as steric repulsion, hydrodynamics, and chemical fields. Yet, these…

Soft Condensed Matter · Physics 2024-06-03 Mario Theers , Elmar Westphal , Kai Qi , Roland G. Winkler , Gerhard Gompper

A striking feature of the collective behavior of spherical microswimmers is that for sufficiently strong self-propulsion they phase-separate into a dense cluster coexisting with a low-density dis- ordered surrounding. Extending our previous…

Soft Condensed Matter · Physics 2016-10-31 Johannes Blaschke , Maurice Maurer , Karthik Menon , Andreas Zöttl , Holger Stark

In this paper we systematically derive a fourth-order continuum theory capable of reproducing mesoscale turbulence in a three-dimensional suspension of microswimmers. We start from overdamped Langevin equations for a generic microscopic…

Soft Condensed Matter · Physics 2018-06-28 Henning Reinken , Sabine H. L. Klapp , Markus Bär , Sebastian Heidenreich

Self-propelled microparticles create flow fields that determine how they interact with surfaces, external flows, and each other. These flow fields fall into distinct classes--pushers, pullers, and neutral swimmers--each exhibiting…

Soft Condensed Matter · Physics 2025-12-25 Lisa Rohde , Gordei Anchutkin , Viktor Holubec , Frank Cichos

Microorganisms ofter move in confined, disordered environments, where hydrodynamic couplings can modify their transport behavior. Using extensive finite-element simulations, we investigate the dynamics of microswimmers -- modeled as…

Soft Condensed Matter · Physics 2026-03-24 Mirko Residori , Sebastian Aland , Christina Kurzthaler

We derive from first principles a three-dimensional theory of self-propelled particle swarming in a viscous fluid environment. Our model predicts emergent collective behavior that depends critically on fluid opacity, mechanism of…

Soft Condensed Matter · Physics 2026-05-12 Yao-Li Chuang , M. R. D'Orsogna , T. Chou

We explore hydrodynamic interactions between microswimmers and corrugated, or rough, surfaces, as found often in biological systems and microfluidic devices. Using the Lorentz reciprocal theorem for viscous flows we derive exact expressions…

Soft Condensed Matter · Physics 2022-08-10 Christina Kurzthaler , Howard A. Stone

We present a two dimensional model of hydrodynamic interaction between a circular swimmer and a circular post at low Reynolds number, using a point singularity description of the swimming activity. We derive a nonlinear dynamical system…

Fluid Dynamics · Physics 2016-12-09 Dario Papavassiliou , Gareth P Alexander

Microswimmers in suspension exhibit collective swimming behaviour, forming various self-organised structures including ordered, aggregated, and turbulent-like structures. When mixed with passive particles phase-separation is known to occur,…

Fluid Dynamics · Physics 2026-03-26 Alexander Chamolly , Takuji Ishikawa

We discuss a micro-swimmer model made of three spheres actuated by an internal active time-periodic force, tied by an elastic potential and submitted to hydrodynamic interactions with thermal noise. The dynamical approach we use, replacing…

Soft Condensed Matter · Physics 2023-11-23 R. Ferretta , R. Di Leonardo , A. Puglisi