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In robot swarms operating under highly restrictive sensing and communication constraints, individuals may need to use direct physical proximity to facilitate information exchange. However, in certain task-related scenarios, this requirement…

Robotics · Computer Science 2019-03-05 Siddharth Mayya , Gennaro Notomista , Dylan Shell , Seth Hutchinson , Magnus Egerstedt

The complex formations exhibited by schooling fish have long been the object of fascination for biologists and physicists. However, the physical and sensory mechanisms leading to organized collective behavior remain elusive. On the physical…

Fluid Dynamics · Physics 2025-07-09 Anand U. Oza , Eva Kanso , Michael J. Shelley

Active particles, which interact hydrodynamically, display a remarkable variety of emergent collective phenomena. We use squirmers to model spherical microswimmers and explore the collective behavior of thousands of them under the influence…

Fluid Dynamics · Physics 2017-10-31 Jan-Timm Kuhr , Johannes Blaschke , Felix Rühle , Holger Stark

Collective behavior of self-propelled particles is observed on a microscale for swimmers such as sperm and bacteria as well as for protein filaments in motility assays. The properties of such systems depend both on their dimensionality and…

Soft Condensed Matter · Physics 2014-01-07 Masoud Abkenar , Kristian Marx , Thorsten Auth , Gerhard Gompper

Microswimmers play an important role in shaping the world around us. The squirmer is a simple model for microswimmer whose cilia oscillations on its spherical surface induce an effective slip velocity to propel itself. The rapid development…

Fluid Dynamics · Physics 2024-11-22 Xinwei Cai , Kuiliang Wang , Gaojin Li , Xin Bian

We confine a dense suspension of motile \textit{Escherichia coli} inside a spherical droplet in a water-in-oil emulsion, creating a "bacterially" propelled droplet. We show that droplets move in a persistent random walk, with a persistence…

Soft Condensed Matter · Physics 2019-12-10 Gabriel Ramos , Maria Luisa Cordero , Rodrigo Soto

The transport properties of colloidal particles in active liquids have been studied extensively. It has led to a deeper understanding of the interactions between passive and active particles. However, the phase behavior of colloidal…

Soft Condensed Matter · Physics 2023-01-30 Pragya Kushwaha , Vivek Semwal , Sayan Maity , Shraddha Mishra , Vijayakumar Chikkadi

In nature, microorganisms must often cope with hostile environmental conditions. To do so they have developed sophisticated cooperative behavior and intricate communication capabilities, such as: direct cell-cell physical interactions via…

Condensed Matter · Physics 2007-05-23 Eshel Ben-Jacob , Inon Cohen , Ido Golding , Yonathan Kozlovsky

We propose a new kind of collective motion where swarms of simple agents are able to find and fix the solution of two-dimensional mazes. The model consists of active memoryless particles interacting exclusively via short-ranged perception…

The interaction between swimming microorganisms or artificial self-propelled colloids and passive (tracer) particles in a fluid leads to enhanced diffusion of the tracers. This enhancement has attracted strong interest, as it could lead to…

Soft Condensed Matter · Physics 2017-07-04 Joost de Graaf , Joakim Stenhammar

The paper presents a two-phase hydrodynamic model for the numerical simulation of collective motion in a thin layer of active colloids containing spherical microswimmers. The model accounts for three fundamental mechanisms governing the…

Soft Condensed Matter · Physics 2025-12-10 A. Kiverin , S. Luguev , I. Yakovenko

Microorganisms are ubiquitous in nature and technology. They inhabit diverse environments ranging from small river tributaries and lakes to oceans, as well as wastewater treatment plants and food manufacturing. In many of these…

Self-propelled particles can exhibit surprising non-equilibrium behaviors, and how they interact with obstacles or boundaries remains an important open problem. Here we show that chemically propelled micro-rods can be captured, with little…

Soft Condensed Matter · Physics 2014-02-21 Daisuke Takagi , Jeremie Palacci , Adam B. Braunschweig , Michael J. Shelley , Jun Zhang

Many active biological particles, such as swimming microorganisms or motor-proteins, do work on their environment by going though a periodic sequence of shapes. Interactions between particles can lead to the phase-synchronization of their…

Soft Condensed Matter · Physics 2022-07-26 Brato Chakrabarti , Michael J. Shelley , Sebastian Fürthauer

Living systems routinely consume energy to achieve motility, often using intricate biomolecular machinery. In this work, we show that active droplets can sustain indefinite self-propulsion of a spherical colloid in an otherwise homogeneous,…

Soft Condensed Matter · Physics 2026-05-01 Benjamin Sorkin , Ned S. Wingreen

In the present work we simulate the basic two-dimensional dynamics of swarming E. coli bacteria on the surface of a moderately soft agar plate. Individual bacteria are modelled by self-propelled ridged bodies (agents), which interact with…

Soft Condensed Matter · Physics 2016-12-31 David Hansmann , Guido Fier , Rubén Carlos Buceta

Typical bodily and environmental fluids encountered by biological swimmers consist of dissolved macromolecules such as proteins and polymers, often rendering them non Newtonian. To mimic such scenarios, we investigate the motion of swimming…

Soft Condensed Matter · Physics 2023-05-23 Prateek Dwivedi , Atishay Shrivastava , Dipin Pillai , Rahul Mangal

Self-propulsion (SP) is a main feature of active particles (AP), such as bacteria or biological micromotors, distinguishing them from passive colloids. A renowned consequence of SP is accumulation at static interfaces, even in the absence…

Soft Condensed Matter · Physics 2017-09-12 U. Marini Bettolo Marconi , A. Sarracino , C. Maggi , A. Puglisi

Adding a non-adsorbing polymer to passive colloids induces an attraction between the particles via the `depletion' mechanism. High enough polymer concentrations lead to phase separation. We combine experiments, theory and simulations to…

Soft Condensed Matter · Physics 2016-07-13 J. Schwarz-Linek , C. Valeriani , A. Cacciuto , M. E. Cates , D. Marenduzzo , A. N. Morozov , W. C. K. Poon

Unicellular microscopic organisms living in aqueous environments outnumber all other creatures on Earth. A large proportion of them are able to self-propel in fluids with a vast diversity of swimming gaits and motility patterns. In this…

Biological Physics · Physics 2021-07-14 Marcos F. Velho Rodrigues , Maciej Lisicki , Eric Lauga