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Recent experiments and numerical studies have drawn attention to the dynamics of active nematics. Two-dimensional active nematics flow spontaneously and exhibit spatiotemporal chaotic flows with proliferation of topological defects in the…

Soft Condensed Matter · Physics 2020-07-08 Farzan Vafa , Mark J. Bowick , M. Cristina Marchetti , Boris I. Shraiman

The dynamics of cellular pattern formation is crucial for understanding embryonic development and tissue morphogenesis. Recent studies have shown that human dermal fibroblasts cultured on liquid crystal elastomers can exhibit an increase in…

Biological Physics · Physics 2023-08-25 Mengyang Gu , Xinyi Fang , Yimin Luo

This essay provides a critical overview of the mathematical kinetic theory of active particles, which is used to model and study collective systems consisting of interacting living entities, such as those involved in behavior and evolution.…

Physics and Society · Physics 2025-10-17 Marina Dolfin , Leone Leonida

Motor proteins drive persistent motion and self-organisation of cytoskeletal filaments. However, state-of-the-art microscopy techniques and continuum modelling approaches focus on large length and time scales. Here, we perform…

Soft Condensed Matter · Physics 2024-06-03 Gerrit Vliegenthart , Arvind Ravichandran , Marisol Ripoll , Thorsten Auth , Gerhard Gompper

Randomly moving active particles can be herded into directed motion by asymmetric geometric structures. Although such a rectification process has been extensively studied due to its fundamental, biological, and technological relevance, a…

Soft Condensed Matter · Physics 2026-03-31 Satyam Anand , Xiaolei Ma , Shuo Guo , Stefano Martiniani , Xiang Cheng

In vitro reconstituted active systems, such as the ATP-driven microtubule bundle suspension developed by the Dogic group, provide a fertile testing ground for elucidating the phenomenology of active liquid crystalline states. Controlling…

Soft Condensed Matter · Physics 2017-01-04 Pau Guillamat , Jordi Ignés-Mullol , Suraj Shankar , M. Cristina Marchetti , Francesc Sagués

Recent studies have shown that packings of cells, both eukaryotic cellular tissues and growing or swarming bacterial colonies, can often be understood as active nematic fluids. A key property of volume-conserving active nematic model…

Soft Condensed Matter · Physics 2022-05-12 Fabian Jan Schwarzendahl , Daniel A. Beller

Active nematics are the nonequilibrium analog of passive liquid crystals in which anisotropic units consume free energy to drive emergent behavior. Similar to liquid crystal (LC) molecules in displays, ordering and dynamics in active…

Quantitative Methods · Quantitative Biology 2022-10-10 Japinder Nijjer , Mrityunjay Kothari , Changhao Li , Thomas Henzel , Qiuting Zhang , Jung-Shen B. Tai , Shuang Zhou , Sulin Zhang , Tal Cohen , Jing Yan

Active matter consists of units that generate mechanical work by consuming energy. Examples include living systems, such as assemblies of bacteria and biological tissues, biopolymers driven by molecular motors, and suspensions of synthetic…

Soft Condensed Matter · Physics 2021-02-04 Song Liu , Suraj Shankar , M. Cristina Marchetti , Yilin Wu

Two-dimensional nonequilibrium nematic steady states, as found in agitated granular-rod monolayers or films of orientable amoeboid cells, were predicted [Europhys. Lett. {\bf 62} (2003) 196] to have giant number fluctuations, with standard…

Soft Condensed Matter · Physics 2009-11-11 Shradha Mishra , Sriram Ramaswamy

Active matter exhibits various forms of non-equilibrium states in the absence of external forcing, including macroscopic steady-state currents. Such states are often too complex to be modelled from first principles and our understanding of…

Soft Condensed Matter · Physics 2020-09-17 Viktor Škultéty , Cesare Nardini , Joakim Stenhammar , Davide Marenduzzo , Alexander Morozov

Self-propelling active matter relies on the conversion of energy from the undirected, nanoscopic scale to directed, macroscopic motion. One of the challenges in the design of synthetic active matter lies in the control of dynamic states, or…

Fluid Dynamics · Physics 2025-03-10 Prashanth Ramesh , Yibo Chen , Petra Räder , Svenja Morsbach , Maziyar Jalaal , Corinna C Maass

Using agent-based simulations of self-propelled particles subject to short-range repulsion and nematic alignment we explore the dynamical phases of a dense active material confined to the surface of a sphere. We map the dynamical phase…

Soft Condensed Matter · Physics 2018-04-25 Silke Henkes , M. Cristina Marchetti , Rastko Sknepnek

Active nematics, in their pure form, have demonstrated a plethora of dynamic and steady state behaviors, including large-scale dynamic structures, collective flows, and intricate multi-spatial temporal dynamics. This complexity further…

Statistical Mechanics · Physics 2024-07-09 Partha Sarathi Mondal , Pawan Kumar Mishra , Tamás Vicsek , Shradha Mishra

We construct the equations of motion for the coupled dynamics of order parameter and concentration for the nematic phase of driven particles on a solid surface, and show that they imply (i) giant number fluctuations, with a standard…

Soft Condensed Matter · Physics 2009-11-07 Sriram Ramaswamy , R. Aditi Simha , John Toner

Active matter comprises individually driven units that convert locally stored energy into mechanical motion. Interactions between driven units lead to a variety of non-equilibrium collective phenomena in active matter. One of such phenomena…

Soft Condensed Matter · Physics 2021-06-15 Mingji Huang , Wensi Hu , Siyuan Yang , Quan-Xing Liu , H. P. Zhang

Swimming bacteria in passive nematics in the form of lyotropic liquid crystals are defined as a new class of active matter known as living liquid crystals in recent studies. It has also been shown that liquid crystal solutions are promising…

Soft Condensed Matter · Physics 2021-09-01 Pranay Bimal Sampat , Shradha Mishra

Active matter classifies systems consisting of self-propelled units which convert the energy stored locally or extracted from their environment into directed motion. It has recently attracted considerable attention due to rich new physics…

Plasma Physics · Physics 2026-04-09 Volodymyr Nosenko

The understanding of dynamics and functioning of biological membranes and in particular of membrane embedded proteins is one of the most fundamental problems and challenges in modern biology and biophysics. In particular the impact of…

Biological Physics · Physics 2009-12-27 Maikel C. Rheinstadter

Planktonic active matter represents an emergent system spanning different scales: individual, population and community; and complexity arising from sub-cellular and cellular to collective and ecosystem scale dynamics. This cross-scale…

Biological Physics · Physics 2023-01-24 Anupam Sengupta