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We report on the dynamics of collective alignment in groups of the cichlid fish, Etroplus suratensis. Focusing on small-to-intermediate sized groups ($10<N<100$), we demonstrate that schooling (highly polarised and coherent motion) is…

Statistical Mechanics · Physics 2020-04-22 Jitesh Jhawar , Richard G. Morris , U. R. Amith-Kumar , M. Danny Raj , Harikrishnan R. , Vishwesha Guttal

Fish rheotaxis, or alignment into flow currents, results from intertwined sensory, neural and actuation mechanisms, all coupled with hydrodynamics to produce a behavior that is critical for upstream migration and position holding in…

Fluid Dynamics · Physics 2016-04-20 Brendan Colvert , Eva Kanso

We experimentally investigate the role of illumination on the collective dynamics of a large school (ca. 50 individuals) of Hemigrammus rhodostomus. The structure of the group, defined using two order parameters, is quantified while…

Adaptation and Self-Organizing Systems · Physics 2023-01-24 Baptiste Lafoux , Jeanne Moscatelli , Ramiro Godoy-Diana , Benjamin Thiria

In open water, social fish gather to form schools, in which fish generally align with each other. In this work, we study how this social behavior evolves when perturbed by artificial obstacles. We measure the collective behavior of a group…

Biological Physics · Physics 2024-07-03 Bruno Ventéjou , Iris Magniez--Papillon , Eric Bertin , Philippe Peyla , Aurélie Dupont

Fish schooling is often modeled with self-propelled particles subject to phenomenological behavioral rules. Although fish are known to sense and exploit flow features, these models usually neglect hydrodynamics. Here, we propose a novel…

Biological Physics · Physics 2018-05-16 Audrey Filella , François Nadal , Clément Sire , Eva Kanso , Christophe Eloy

Schooling, an archetype of collective behavior, emerges from the interactions of fish responding to visual and other informative cues mediated by their aqueous environment. In this context, a fundamental and largely unexplored question…

The complex formations exhibited by schooling fish have long been the object of fascination for biologists and physicists. However, the physical and sensory mechanisms leading to organized collective behavior remain elusive. On the physical…

Fluid Dynamics · Physics 2025-07-09 Anand U. Oza , Eva Kanso , Michael J. Shelley

Fish schools are ubiquitous in marine life. Although flow interactions are thought to be beneficial for schooling, their exact effects on the speed, energetics, and stability of the group remain elusive. Recent experiments suggest that flow…

Fluid Dynamics · Physics 2021-08-04 Sina Heydari , Eva Kanso

There is increasing evidence that fish gain energetic benefits when they swim in a school. The most recent indications of such benefits are a lower tail (or fin) beat at the back of a school and reduced oxygen consumption in schooling fish…

Quantitative Methods · Quantitative Biology 2013-07-30 C. K. Hemelrijk , D. A. P. Reid , H. Hildenbrandt , J. T. Padding

A novel model of the wake of swimming fish is developed and incorporated into a dynamical model of a fish school to explore the effect of hydrodynamics on the emergent behavior in schooling fish. The model incorporates well-established…

Fluid Dynamics · Physics 2025-07-11 Ji Zhou , Jung-Hee Seo , Rajat Mittal

Fish schools are able to display a rich variety of collective states and behavioural responses when they are confronted to threats. However a school's response to perturbations may be different depending on its collective state. Here we use…

Quantitative Methods · Quantitative Biology 2015-02-04 Daniel S. Calovi , Ugo Lopez , Paul Schuhmacher , Hugues Chaté , Clément Sire , Guy Theraulaz

Fish, birds, insects and robots frequently swim or fly in groups. During their 3 dimensional collective motion, these agents do not stop, they avoid collisions by strong short-range repulsion, and achieve group cohesion by weak long-range…

Soft Condensed Matter · Physics 2018-07-04 Illes J. Farkas , Shuo-Hong Wang

Marine plankton are usually modeled as settling elongated micro-swimmers. For the first time, we consider the torque induced by fluid inertia on such swimmers, and we discover that they spontaneously swim in the direction opposite to…

Biological Physics · Physics 2022-05-12 Jingran Qiu , Zhiwen Cui , Eric Climent , Lihao Zhao

Schooling fish often self-organize into a variety of collective patterns, from polarized schooling to rotational milling. Mathematical models support the emergence of these large-scale patterns from local decentralized interactions, in the…

Adaptation and Self-Organizing Systems · Physics 2025-07-23 Alyssa Chan , Eva Kanso

One of the most highly debated questions in the field of animal swarming and social behaviour, is the collective random patterns and chaotic behaviour formed by some animal species, in particular if there is a danger. Is such a behaviour…

Populations and Evolution · Quantitative Biology 2017-01-04 Usama Kadri , Franz Brümmer , Anan Kadri

The collective motion of self-driven agents is a phenomenon of great interest in interacting particle systems. In this paper, we develop and analyze a model of agent motion in one dimension with periodic boundaries using a stochastic…

Dynamical Systems · Mathematics 2023-03-28 Matthias Ehrhardt , Thomas Kruse , Antoine Tordeux

While a rich variety of self-propelled particle models propose to explain the collective motion of fish and other animals, rigorous statistical comparison between models and data remains a challenge. Plausible models should be flexible…

Quantitative Methods · Quantitative Biology 2017-04-28 Maksym Romenskyy , James E. Herbert-Read , Ashley J. W. Ward , David J. T. Sumpter

We combine extensive data analyses with a modeling approach to measure, disentangle, and reconstruct the actual functional form of interactions involved in the coordination of swimming in Rummy-nose tetra (Hemigrammus rhodostomus). This…

Waves and oscillations are commonly observed in the dynamics of self-driven agents such as pedestrians or vehicles. Interestingly, many factors may perturb the stability of space homogeneous streaming, leading to the spontaneous formation…

Physics and Society · Physics 2021-06-10 Martin Friesen , Hanno Gottschalk , Barbara Rüdiger , Antoine Tordeux

We present new constrained and free-swimming experiments and simulations of a pair of pitching hydrofoils interacting in a minimal school. The hydrofoils have an out-of-phase synchronization and they are varied through in-line, staggered,…

Fluid Dynamics · Physics 2023-08-01 Pedro Costa Ormonde , Melike Kurt , Amin Mivehchi , Keith W. Moored
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