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Phylogenetic networks play an important role in evolutionary biology as, other than phylogenetic trees, they can be used to accommodate reticulate evolutionary events such as horizontal gene transfer and hybridization. Recent research has…

Populations and Evolution · Quantitative Biology 2024-03-19 Linda Knüver , Mareike Fischer , Marc Hellmuth , Kristina Wicke

Several indices that measure the degree of balance of a rooted phylogenetic tree have been proposed so far in the literature. In this work we define and study a new index of this kind, which we call the total cophenetic index: the sum, over…

Populations and Evolution · Quantitative Biology 2014-02-11 Arnau Mir , Francesc Rossello , Lucia Rotger

Tree balance plays an important role in different research areas like theoretical computer science and mathematical phylogenetics. For example, it has long been known that under the Yule model, a pure birth process, imbalanced trees are…

Populations and Evolution · Quantitative Biology 2020-12-18 Mareike Fischer

The Sackin and Colless indices are two widely-used metrics for measuring the balance of trees and for testing evolutionary models in phylogenetics. This short paper contributes two results about the Sackin and Colless indices of trees. One…

Populations and Evolution · Quantitative Biology 2022-07-20 Gary Goh , Michael Fuchs , Louxin Zhang

The study of species organization and their clustering by genetic or phenotypic similarity is carried out with the tools of phylogenetic trees. An important structural property of phylogenetic trees is the balance, which measures how taxa…

Populations and Evolution · Quantitative Biology 2020-09-01 T. Araújo Lima , Flavia M. D. Marquitti , Marcus A. M. de Aguiar

Measures of tree balance play an important role in various research areas, for example in phylogenetics. There they are for instance used to test whether an observed phylogenetic tree differs significantly from a tree generated by the Yule…

Combinatorics · Mathematics 2019-07-23 Mareike Fischer , Lina Herbst , Kristina Wicke

We solve a class of optimization problems for (phylogenetic) $X$-trees or their shapes. These problems have recently appeared in different contexts, e.g. in the context of the impact of tree shapes on the size of TBR neighborhoods, but so…

Populations and Evolution · Quantitative Biology 2015-10-28 Mareike Fischer , Volkmar Liebscher

We define a new balance index for rooted phylogenetic trees based on the symmetry of the evolutive history of every set of 4 leaves. This index makes sense for multifurcating trees and it can be computed in time linear in the number of…

Populations and Evolution · Quantitative Biology 2019-03-25 Tomás M. Coronado , Arnau Mir , Francesc Rosselló , Gabriel Valiente

Measures of tree balance play an important role in different research areas such as mathematical phylogenetics or theoretical computer science. The balance of a tree is usually quantified in a single number, called a balance or imbalance…

Combinatorics · Mathematics 2024-06-28 Bryan Currie , Kristina Wicke

Tree balance plays an important role in various research areas in phylogenetics and computer science. Typically, it is measured with the help of a balance index or imbalance index. There are more than 25 such indices available, recently…

Populations and Evolution · Quantitative Biology 2025-08-20 Sean Cleary , Mareike Fischer , Katherine St. John

Tree shape statistics, particularly measures of tree (im)balance, play an important role in the analysis of the shape of phylogenetic trees. With applications ranging from testing evolutionary models to studying the impact of fertility…

Populations and Evolution · Quantitative Biology 2024-10-15 Sophie J. Kersting , Kristina Wicke , Mareike Fischer

Phylogenetic tree shapes capture fundamental signatures of evolution. We consider ``ranked'' tree shapes, which are equipped with a total order on the internal nodes compatible with the tree graph. Recent work has established an elegant…

Populations and Evolution · Quantitative Biology 2026-03-10 Chris Jennings-Shaffer , Ziyue , Chen , Julia A Palacios , Frederick A Matsen

Phylogenetic trees capture evolutionary relationships among species and reflect the forces that shaped them. While many studies rely on branch length information, the topology of phylogenetic trees (particularly their degree of imbalance)…

Populations and Evolution · Quantitative Biology 2025-10-16 Stephan Kleinbölting , Nigel Goldenfeld , Johannes Berg

The Colless index for bifurcating phylogenetic trees, introduced by Colless (1982), is defined as the sum, over all internal nodes $v$ of the tree, of the absolute value of the difference of the sizes of the clades defined by the children…

Populations and Evolution · Quantitative Biology 2020-07-30 Tomás M. Coronado , Arnau Mir , Francesc Rosselló

The Sackin index is an important measure for the balance of phylogenetic trees. We investigate two extensions of the Sackin index to the class of galled trees and two of its subclasses (simplex galled trees and normal galled trees) where we…

Populations and Evolution · Quantitative Biology 2025-02-21 Michael Fuchs , Bernhard Gittenberger

One of the main applications of balance indices is in tests of null models of evolutionary processes. The knowledge of an exact formula for a statistic of a balance index, holding for any number n of leaves, is necessary in order to use…

Populations and Evolution · Quantitative Biology 2012-10-22 Gabriel Cardona , Arnau Mir , Francesc Rossello

We investigate the structure of trees that have greatest maximum eigenvalue among all trees with a given degree sequence. We show that in such an extremal tree the degree sequence is non-increasing with respect to an ordering of the…

Combinatorics · Mathematics 2008-04-18 Tuerker Biyikoglu , Marc Hellmuth , Josef Leydold

There are several common ways to encode a tree as a matrix, such as the adjacency matrix, the Laplacian matrix (that is, the infinitesimal generator of the natural random walk), and the matrix of pairwise distances between leaves. Such…

Populations and Evolution · Quantitative Biology 2007-05-23 Frederick A. Matsen , Steven N. Evans

Full binary trees naturally represent commutative non-associative products. There are many important examples of these products: finite-precision floating-point addition and NAND gates, among others. Balance in such a tree is highly…

Discrete Mathematics · Computer Science 2021-08-27 Laura Monroe

Bayesian inference is now a leading technique for reconstructing phylogenetic trees from aligned sequence data. In this short note, we formally show that the maximum posterior tree topology provides a statistically consistent estimate of a…

Populations and Evolution · Quantitative Biology 2013-07-12 Mike Steel
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