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Although the analysis of rooted tree shape has wide-ranging applications, notions of tree balance have developed independently in different domains. In computer science, a balanced tree is one that enables efficient updating and retrieval…

Quantitative Methods · Quantitative Biology 2025-07-14 Veselin Manojlović , Armaan Ahmed , Yannick Viossat , Robert Noble

Tree balance plays an important role in different research areas like theoretical computer science and mathematical phylogenetics. For example, it has long been known that under the Yule model, a pure birth process, imbalanced trees are…

Populations and Evolution · Quantitative Biology 2020-12-18 Mareike Fischer

Phylogenetic diversity indices are commonly used to rank the elements in a collection of species or populations for conservation purposes. The derivation of these indices is typically based on some quantitative description of the…

Populations and Evolution · Quantitative Biology 2024-03-26 Vincent Moulton , Andreas Spillner , Kristina Wicke

One of the main applications of balance indices is in tests of null models of evolutionary processes. The knowledge of an exact formula for a statistic of a balance index, holding for any number n of leaves, is necessary in order to use…

Populations and Evolution · Quantitative Biology 2012-10-22 Gabriel Cardona , Arnau Mir , Francesc Rossello

Tree balance plays an important role in various research areas in phylogenetics and computer science. Typically, it is measured with the help of a balance index or imbalance index. There are more than 25 such indices available, recently…

Populations and Evolution · Quantitative Biology 2025-08-20 Sean Cleary , Mareike Fischer , Katherine St. John

The Sackin index is an important measure for the balance of phylogenetic trees. We investigate two extensions of the Sackin index to the class of galled trees and two of its subclasses (simplex galled trees and normal galled trees) where we…

Populations and Evolution · Quantitative Biology 2025-02-21 Michael Fuchs , Bernhard Gittenberger

We introduce a scale-free method for testing the proportionality of branch lengths between two phylogenetic trees that have the same topology and contain the same set of taxa. This method scales both trees to a total length of 1 and sums up…

Quantitative Methods · Quantitative Biology 2015-03-16 Yichen Zheng , William Ott , Chinmaya Gupta , Dan Graur

Measures of tree balance play an important role in many different research areas such as mathematical phylogenetics or theoretical computer science. Typically, tree balance is quantified by a single number which is assigned to the tree by a…

Populations and Evolution · Quantitative Biology 2025-07-31 Mareike Fischer , Tom Niklas Hamann , Kristina Wicke

Phylogenetic trees capture evolutionary relationships among species and reflect the forces that shaped them. While many studies rely on branch length information, the topology of phylogenetic trees (particularly their degree of imbalance)…

Populations and Evolution · Quantitative Biology 2025-10-16 Stephan Kleinbölting , Nigel Goldenfeld , Johannes Berg

We introduce a cluster evaluation technique called Tree Index. Our Tree Index algorithm aims at describing the structural information of the clustering rather than the quantitative format of cluster-quality indexes (where the representation…

Machine Learning · Computer Science 2020-03-25 A. H. Beg , Md Zahidul Islam , Vladimir Estivill-Castro

The study of species organization and their clustering by genetic or phenotypic similarity is carried out with the tools of phylogenetic trees. An important structural property of phylogenetic trees is the balance, which measures how taxa…

Populations and Evolution · Quantitative Biology 2020-09-01 T. Araújo Lima , Flavia M. D. Marquitti , Marcus A. M. de Aguiar

A phylogenetic tree is an edge-weighted binary tree, with leaves labelled by a collection of species, that represents the evolutionary relationships between those species. For such a tree, a phylogenetic diversity index is a function that…

Optimization and Control · Mathematics 2026-05-08 Martin Frohn , Kerry Manson

Effects like selection in evolution as well as fertility inheritance in the development of populations can lead to a higher degree of asymmetry in evolutionary trees than expected under a null hypothesis. To identify and quantify such…

Populations and Evolution · Quantitative Biology 2021-08-05 Sophie J. Kersting , Mareike Fischer

Planning for the protection of species often involves difficult choices about which species to prioritize, given constrained resources. One way of prioritizing species is to consider their "evolutionary distinctiveness", i.e. their relative…

Populations and Evolution · Quantitative Biology 2021-07-05 Mareike Fischer , Andrew Francis , Kristina Wicke

As researchers collect increasingly large molecular data sets to reconstruct the Tree of Life, the heterogeneity of signals in the genomes of diverse organisms poses challenges for traditional phylogenetic analysis. A class of phylogenetic…

Populations and Evolution · Quantitative Biology 2015-09-11 Liang Liu , Zhenxiang Xi , Shaoyuan Wu , Charles Davis , Scott V. Edwards

Selective inference is considered for testing trees and edges in phylogenetic tree selection from molecular sequences. This improves the previously proposed approximately unbiased test by adjusting the selection bias when testing many trees…

Applications · Statistics 2019-05-27 Hidetoshi Shimodaira , Yoshikazu Terada

In phylogenetic networks, it is desirable to estimate edge lengths in substitutions per site or calendar time. Yet, there is a lack of scalable methods that provide such estimates. Here we consider the problem of obtaining edge length…

Populations and Evolution · Quantitative Biology 2024-08-06 Jingcheng Xu , Cécile Ané

The algebraic properties of flattenings and subflattenings provide direct methods for identifying edges in the true phylogeny -- and by extension the complete tree -- using pattern counts from a sequence alignment. The relatively small…

Populations and Evolution · Quantitative Biology 2022-05-06 Joshua Stevenson , Barbara Holland , Michael Charleston , Jeremy Sumner

Comparative analyses of phylogenetic trees typically require identical taxon sets, however, in practice, trees often include distinct but overlapping taxa. Pruning non-shared leaves discards phylogenetic signal, whereas tree completion can…

Populations and Evolution · Quantitative Biology 2026-04-28 Aleksandr Koshkarov , Nadia Tahiri

The Shapley Value and the Fair Proportion Index of phylogenetic trees have been frequently discussed as prioritization tools in conservation biology. Both indices rank species according to their contribution to total phylogenetic diversity,…

Populations and Evolution · Quantitative Biology 2017-07-28 Kristina Wicke , Mareike Fischer