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In this article we derive and test the fluctuating hydrodynamic description of active particles interacting via taxis and quorum sensing, both for mono-disperse systems and for mixtures of co-existing species of active particles. We compute…

Statistical Mechanics · Physics 2025-01-14 Alberto Dinelli , Jérémy O'Byrne , Julien Tailleur

A simple theoretical approach is used to investigate active colloids at the free interface and near repulsive substrates. We employ dynamical density functional theory to determine the steady-state density profiles in an effective…

Soft Condensed Matter · Physics 2017-02-03 René Wittmann , Joseph M. Brader

The classical macroscopic chemotaxis equations have previously been derived from an individual-based description of the tactic response of cells that use a "run-and-tumble" strategy in response to environmental cues. Here we derive…

Cell Behavior · Quantitative Biology 2007-05-23 Radek Erban , Hans G. Othmer

Chemotaxis is the response of a particle to a gradient in the chemical composition of the environment. While it was originally observed for biological organisms, it is of great interest in the context of synthetic active particles such as…

Soft Condensed Matter · Physics 2017-05-02 Laurens Deprez , Pierre de Buyl

In colloidal suspensions, the competition between attractive and repulsive interactions gives rise to a rich and complex phenomenology. Here, we study the equilibrium phase diagram of a model system using a DLVO interaction potential by…

Soft Condensed Matter · Physics 2009-11-11 A. de Candia , E. Del Gado , A. Fierro , N. Sator , M. Tarzia , A. Coniglio

Active particles with their characteristic feature of self-propulsion are regarded as the simplest models for motility in living systems. The accumulation of active particles in low activity regions has led to the general belief that…

Soft Condensed Matter · Physics 2021-05-26 Hidde D. Vuijk , Holger Merlitz , Michael Lang , Abhinav Sharma , Jens-Uwe Sommer

Swimmers and self-propelled particles are physical models for the collective behaviour and motility of a wide variety of living systems, such as bacteria colonies, bird flocks and fish schools. Such artificial active materials are amenable…

Microphase separation is common in active biological systems as exemplified by the separation of RNA and DNA-rich phases in the cell nucleus driven by the transcriptional activity of polymerase enzymes acting similarly to amphiphiles in a…

Soft Condensed Matter · Physics 2025-07-22 Rakesh Chatterjee , Hui-Shun Kuan , Frank Julicher , Vasily Zaburdaev

Many fascinating properties of biological active matter crucially depend on the capacity of constituting entities to perform directed motion, e.g., molecular motors transporting vesicles inside cells or bacteria searching for food. While…

Statistical Mechanics · Physics 2023-06-21 Pietro Luigi Muzzeddu , Édgar Roldán , Andrea Gambassi , Abhinav Sharma

We consider a rigid assembly of two active Brownian particles, forming an active colloidal dimer, in a gradient of activity. We show analytically that depending on the relative orientation of the two particles the active dimer accumulates…

Soft Condensed Matter · Physics 2022-08-17 Hidde D. Vuijk , Sophie Klempahn , Holger Merlitz , Jens-Uwe Sommer , Abhinav Sharma

Biomolecular condensates play a central role in the spatial organization of living matter. Their formation is now well understood as a form of liquid-liquid phase separation that occurs very far from equilibrium. For instance, they can be…

Soft Condensed Matter · Physics 2024-10-14 Roxanne Berthin , Jacques Fries , Marie Jardat , Vincent Dahirel , Pierre Illien

Phase separating systems that are maintained away from thermodynamic equilibrium via molecular processes represent a class of active systems, which we call active emulsions. These systems are driven by external energy input for example…

Soft Condensed Matter · Physics 2019-06-07 Christoph A. Weber , David Zwicker , Frank Jülicher , Chiu Fan Lee

Many biological processes are supported by special molecules, called motor proteins or molecular motors, that transport cellular cargoes along linear protein filaments and can reversibly associate to their tracks. Stimulated by these…

Statistical Mechanics · Physics 2021-11-17 Akriti Jindal , Anatoly B. Kolomeisky , Arvind Kumar Gupta

We consider a mixture of passive (i.e., Brownian) and active (e.g., bacterial or colloidal swimmers) particles, and analyze the stability conditions of either uniformly mixed or phase segregated steady states consisting of phases enriched…

Statistical Mechanics · Physics 2015-09-21 Alexander Y. Grosberg , Jean-François Joanny

The transport properties of colloidal particles in active liquids have been studied extensively. It has led to a deeper understanding of the interactions between passive and active particles. However, the phase behavior of colloidal…

Soft Condensed Matter · Physics 2023-01-30 Pragya Kushwaha , Vivek Semwal , Sayan Maity , Shraddha Mishra , Vijayakumar Chikkadi

By means of classical density functional theory and its dynamical extension, we consider a colloidal fluid with spherically-symmetric competing interactions, which are well known to exhibit a rich bulk phase behavior. This includes complex…

Soft Condensed Matter · Physics 2018-06-11 Daniel Stopper , Roland Roth

Active filaments, such as microtubules with attached cargo-carrying motor proteins, are important dynamic structures for fluid transport in and around living cells. The mathematical models of active filaments appearing in the literature…

Fluid Dynamics · Physics 2025-08-19 Ilteber R. Ozdemir , Bethany Clarke , Yongyun Hwang , Eric E. Keaveny

Systems containing active components are intrinsically out of equilibrium, while binary mixtures reach their equilibrium configuration when complete phase separation is achieved. Active particles are found to stabilise non-equilibrium…

Soft Condensed Matter · Physics 2025-03-11 Javier Diaz , Ignacio Pagonabarraga

Aligning self-propelled particles undergo a nonequilibrium flocking transition from apolar to polar phases as their interactions become stronger. We propose a thermodynamically consistent lattice model, in which the internal state of the…

Statistical Mechanics · Physics 2025-08-08 Karel Proesmans , Gianmaria Falasco , Atul Tanaji Mohite , Massimiliano Esposito , Étienne Fodor

To study the impact of active systems on their surroundings, we introduce a model that couples an active nematic fluid to an isotropic substrate fluid via friction. We numerically show that as the active layer develops turbulence, the…

Soft Condensed Matter · Physics 2025-12-01 Gianmarco Spera , Julia M. Yeomans , Sumesh P. Thampi
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