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We study the mechanical stiffening behavior in two-dimensional (2D) cross-linked networks of semiflexible biopolymer filaments under simple shear. Filamental constituents immersed in a fluid undergo thermally excited bending motions.…

Biological Physics · Physics 2007-05-23 T. Van Dillen , P. R. Onck , E. Van der Giessen

The mechanical properties of a thin, planar material, perfused by an embedded flow network, can be changed locally and globally by the fluid transport and storage, resulting in small or large-scale deformation, such as out-of-plane…

Soft Condensed Matter · Physics 2023-03-15 Yongtian Luo , Che-Ling Ho , Brent R. Helliker , Eleni Katifori

We address the folding induced by differential growth in soft layered solids via an elementary model that consists of a soft growing neo-Hookean elastic layer adhered to a deep elastic substrate. As the layer/substrate modulus ratio is…

Soft Condensed Matter · Physics 2015-09-30 Tuomas Tallinen , John S. Biggins

Surface tension is a prominent factor for the deformation of solids at micro-/nano-scale. This paper investigates the effects of surface tension on the two-dimensional contact problems of an elastic layer bonded to the rigid substrate.…

Soft Condensed Matter · Physics 2018-10-29 Weike Yuan , Gangfeng Wang

Rigidity is an emergent property of materials - it is not a feature of individual components that comprise the structure, but instead arises from interactions between many constituent parts. Recently, it has been recognized that…

Soft Condensed Matter · Physics 2025-08-27 Kelly Aspinwall , Tyler Hain , M. Lisa Manning

One of the hallmarks of pre-migratory tumors is the progressive loss of compact morphology. To investigate how tumors may intrinsically regulate their shape during growth, we employ a three-dimensional (3D) vertex model of multicellular…

Soft Condensed Matter · Physics 2026-02-09 Urban Železnik , Matej Krajnc , Tanmoy Sarkar

Dispersions of immiscible liquids, such as emulsions and polymer blends, are at the core of many industrial applications which makes the understanding of their properties (morphology, stability, etc.) of great interest. A wide range of…

Chemical Physics · Physics 2017-08-10 Anaïs Giustiniani , Wiebke Drenckhan , Christophe Poulard

Inspired by the ubiquity of composite filamentous networks in nature we investigate models of biopolymer networks that consist of interconnected floppy and stiff filaments. Numerical simulations carried out in three dimensions allow us to…

Biomolecules · Quantitative Biology 2015-05-19 Elisabeth M. Huisman , Claus Heussinger , Cornelis Storm , Gerard T. Barkema

Many biological tissues feature a heterogeneous network of fibers whose tensile and bending rigidity contribute substantially to these tissues' elastic properties. Rigidity percolation has emerged as a important paradigm for relating these…

The non-linear stress-strain relation for crosslinked polymer networks is studied using molecular dynamics simulations. Previously we demonstrated the importance of trapped entanglements in determining the elastic and relaxational…

Soft Condensed Matter · Physics 2009-10-31 Gary S. Grest , Mathias Puetz , Kurt Kremer , Ralf Everaers

Many of the cell membrane vital functions are achieved by the self-organization of the proteins and biopolymers embedded in it. The protein dynamics are in part determined by its drag. A large number of these proteins can polymerize to form…

Fluid Dynamics · Physics 2023-09-13 Wenzheng Shi , Moslem Moradi , Ehssan Nazockdast

We propose a mechanism for mechanical regulation at the membrane of living cells, based on the exchange of membrane area between the cell membrane and a membrane reservoir. The reservoir is composed of invaginated membrane microdomains…

Subcellular Processes · Quantitative Biology 2007-05-23 Pierre Sens , Matthew S. Turner

We present a continuum model of the coupling between cells and substrate that accounts for some of the observed substrate-stiffness dependence of cell properties. The cell is modeled as an elastic active gel, adapting recently developed…

Cell Behavior · Quantitative Biology 2015-05-28 Shiladitya Banerjee , M. Cristina Marchetti

Networks of stiff fibers govern the elasticity of biological structures such as the extracellular matrix of collagen. These networks are known to stiffen nonlinearly under shear or extensional strain. Recently, it has been shown that such…

Soft Condensed Matter · Physics 2024-02-01 Sihan Chen , Tomer Markovich , Fred C. MacKintosh

The structure of polymers at solid interfaces evolves over time, but the corresponding changes in their rheological properties remain poorly understood. Here, using a home-built quartz tuning fork atomic force microscope-based…

Fluid Dynamics · Physics 2025-08-22 Jaewon Shim , Manhee Lee , Wonho Jhe

The ability of cells to sense and respond to the mechanical properties of their environments is fundamental to a range of cellular behaviours, with substrate stiffness increasingly being found to be a key signalling factor. Although active…

Cell Behavior · Quantitative Biology 2019-09-04 Carina M. Dunlop

Disordered fibrous networks are ubiquitous in nature as major structural components of living cells and tissues. The mechanical stability of networks generally depends on the degree of connectivity: only when the average number of…

Biological Physics · Physics 2016-06-22 A. Sharma , A. J. Licup , R. Rens , M. Sheinman , K. A. Jansen , G. H. Koenderink , F. C. MacKintosh

Biological cells in soft materials can be modeled as anisotropic force contraction dipoles. The corresponding elastic interaction potentials are long-ranged ($\sim 1/r^3$ with distance $r$) and depend sensitively on elastic constants,…

Soft Condensed Matter · Physics 2009-11-07 U. S. Schwarz , S. A. Safran

The mechanical properties of the cell depend crucially on the tension of its cytoskeleton, a biopolymer network that is put under stress by active motor proteins. While the fibrous nature of the network is known to strongly affect the…

Soft Condensed Matter · Physics 2018-05-15 Pierre Ronceray , Chase P. Broedersz , Martin Lenz

We present a theory for the elasticity of cross-linked stiff polymer networks. Stiff polymers, unlike their flexible counterparts, are highly anisotropic elastic objects. Similar to mechanical beams stiff polymers easily deform in bending,…

Soft Condensed Matter · Physics 2007-11-26 C. Heussinger , B. Schaefer , E. Frey
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