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Classical elasticity is concerned with bodies that can be modeled as smooth manifolds endowed with a reference metric that represents local equilibrium distances between neighboring material elements. The elastic energy associated with a…

Materials Science · Physics 2015-06-30 Raz Kupferman , Michael Moshe , Jake P. Solomon

We show that back-flow, the coupling between the order parameter and the velocity fields, has a significant effect on the motion of defects in nematic liquid crystals. In particular the defect speed can depend strongly on the topological…

Soft Condensed Matter · Physics 2007-05-23 Geza Toth , Colin Denniston , Julia M. Yeomans

Topological defects are a ubiquitous phenomenon in diverse physical systems. In nematic liquid crystals (LCs), they are dynamic, physicochemically distinct, sensitive to stimuli, and are thereby promising for a range of applications.…

Soft Condensed Matter · Physics 2023-12-25 Shengzhu Yi , Hao Chen , Xinyu Wang , Miao Jiang , Bo Li , Qi-huo Wei , Rui Zhang

This study theoretically investigates how anisotropic curved membrane components (CMCs) control vesicle morphology through curvature sensing, nematic alignment, topological defects, and volume constraints. By comparing arc-shaped and…

Soft Condensed Matter · Physics 2025-06-18 Yoav Ravid , Samo Penič , Luka Mesarec , Nir S. Gov , Veronika Kralj-Iglič , Aleš Iglič , Mitja Drab

In cell extrusion, a cell embedded in an epithelial monolayer loses its apical or basal surface and is subsequently squeezed out of the monolayer by neighboring cells. Cell extrusions occur during apoptosis, epithelial-mesenchymal…

Cell Behavior · Quantitative Biology 2020-04-24 Satoru Okuda , Koichi Fujimoto

Topological defects in active liquid crystals can be confined by introducing gradients of activity. Here, we examine the dynamical behavior of two defects confined by a sharp gradient of activity that separates an active circular region and…

Soft Condensed Matter · Physics 2021-06-16 Ali Mozaffari , Rui Zhang , Noe Atzin , Juan J. de Pablo

Coherent angular rotation of epithelial cells is thought to contribute to many vital physiological processes including tissue morphogenesis and glandular formation. However, factors regulating this motion, and the implications of this…

Cell Behavior · Quantitative Biology 2016-05-10 S. S. Soumya , Animesh Gupta , Andrea Cugno , Luca Deseri , Kaushik Dayal , Dibyendu Das , Shamik Sen , Mandar M. Inamdar

Cell monolayers are a central model system to tissue biophysics. In vivo, epithelial tissues are curved on the scale of microns, and curvature's role in the onset of spontaneous tissue flows is still not well-understood. Here, we present a…

Biological Physics · Physics 2022-09-21 Samuel Bell , Shao-Zhen Lin , Jean-François Rupprecht , Jacques Prost

We characterise the particlelike kinematics of charge-carrying topological defects in nematic media via a geometric field theory. This differs from the theory of electromagnetism, with which it is often compared, due to the absence of…

Soft Condensed Matter · Physics 2026-05-22 Joseph Pollard , Richard G. Morris

We investigate experimentally and numerically the defect configurations emerging when a cholesteric liquid crystal is confined to a spherical shell. We uncover a rich scenario of defect configurations, some of them non-existent in nematic…

Soft Condensed Matter · Physics 2017-09-13 Alexandre Darmon , Michael Benzaquen , Simon Čopar , Olivier Dauchot , Teresa Lopez-Leon

Cell layers are often categorized as contractile or extensile active nematics but recent experiments on neural progenitor cells with induced $+1$ topological defects challenge this classification. In a bottom-up approach, we first study a…

Soft Condensed Matter · Physics 2024-09-13 Zihui Zhao , Yisong Yao , He Li , Yongfeng Zhao , Yujia Wang , Hepeng Zhang , Hugues Chat'e , Masaki Sano

Topological defects play a key role in two-dimensional active nematics, and a transient role in two-dimensional active polar fluids. In this paper, we study both the transient and long-time behavior of defects in two-dimensional active…

Soft Condensed Matter · Physics 2020-09-24 Farzan Vafa

Polar active matter - including animal herds, aggregates of motile cells and active colloids - often forms coordinated migration patterns, such as flocking. This orderly motion can be disrupted by full-integer topological defects…

Soft Condensed Matter · Physics 2026-05-26 Luiza Angheluta , Anna Lång , Emma Lång , Stig Ove Bøe

A rich variety of specific multidomain textures recently observed in antiferromagnetically coupled multilayers with perpendicular anisotropy include regular (equilibrium) multidomain states as well as different types of topological magnetic…

Mesoscale and Nanoscale Physics · Physics 2009-11-13 N. S. Kiselev , U. K. Roessler , A. N. Bogdanov , O. Hellwig

Epithelial cell monolayers exhibit traveling mechanical waves. We rationalize this observation thanks to a hydrodynamic description of the monolayer as a compressible, active and polar material. We show that propagating waves of the cell…

Biological Physics · Physics 2018-10-01 Shunsuke Yabunaka , Philippe Marcq

We study long-range morphological changes in atomic monolayers on solid substrates induced by different types of defects; e.g., by monoatomic steps in the surface, or by the tip of an atomic force microscope (AFM), placed at some distance…

Soft Condensed Matter · Physics 2009-11-07 H. Schiessel , G. Oshanin , A. M. Cazabat , M. Moreau

Active matter is characterized by its ability to induce motion by self-generated stress. In the case of a solid, such motion can lead to shape transformations. The stress-generating components can be anisotropic endowing the material with…

Soft Condensed Matter · Physics 2022-05-16 D. J. G. Pearce , S. Gat , G. Livne , A. Bernheim-Groswasser , K. Kruse

To enhance the understanding of the behavior of active nematic, it is important to understand the behavior of topological defects. In this paper, we study the configuration of topological defects of a two-dimensional active nematic around a…

Soft Condensed Matter · Physics 2026-02-16 Hiroki Matsukiyo , Jun-ichi Fukuda

We investigate the geometrical and mechanical properties of adherent cells characterized by a highly anisotropic actin cytoskeleton. Using a combination of theoretical work and experiments on micropillar arrays, we demonstrate that the…

Cultured stem cells have become a standard platform not only for regenerative medicine and developmental biology but also for biophysical studies. Yet, the characterization of cultured stem cells at the level of morphology and macroscopic…

Soft Condensed Matter · Physics 2022-06-23 Kyogo Kawaguchi , Ryoichiro Kageyama , Masaki Sano
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