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It is now widely recognized that mechanical interactions between cells play a crucial role in epithelial morphogenesis, yet understanding the mechanisms through which stress and deformation affect cell behavior remains an open problem due…

Tissues and Organs · Quantitative Biology 2017-01-20 Nicholas Noll , Madhav Mani , Idse Heemskerk , Sebastian Streichan , Boris I. Shraiman

Morphogenesis, tissue regeneration and cancer invasion involve transitions in tissue morphology. These transitions, caused by collective cell migration (CCM), have been interpreted as active wetting/de-wetting transitions. This phenomenon…

Cell Behavior · Quantitative Biology 2022-08-29 Ivana Pajic-Lijakovic , Milan Milivojevic

Experiments and theory have shown that cell monolayers and epithelial tissues exhibit solid-liquid and glass-liquid transitions. These transitions are biologically relevant to our understanding of embryonic development, wound healing, and…

Soft Condensed Matter · Physics 2020-07-22 Benjamin Loewe , Michael Chiang , Davide Marenduzzo , M. Cristina Marchetti

Deformations of cell sheets during morphogenesis are driven by developmental processes such as cell division and cell shape changes. In morphoelastic shell theories of development, these processes appear as variations of the intrinsic…

Soft Condensed Matter · Physics 2021-03-03 Pierre A. Haas , Raymond E. Goldstein

A theoretical model for stratified epithelium is presented. The viscoelastic properties of the tissue is assumed to be dependent on the spatial distribution of proliferative and differentiated cells. Based on this assumption, a hydrodynamic…

Biological Physics · Physics 2016-06-08 Wei-Ting Yeh , Hsuan-Yi Chen

In proliferating epithelia of mammalian skin, cells of irregular polygonal-like shapes pack into complex nearly flat two-dimensional structures that are pliable to deformations. In this work, we employ various sensitive correlation…

Statistical Mechanics · Physics 2016-12-07 Duyu Chen , Wen Yih Aw , Danelle Devenport , Salvatore Torquato

Embryonic tissues deform across broad spatial and temporal scales and relax stress through active rearrangements. A quantitative link between cell-scale activity, spatial forcing, and emergent tissue-scale mechanics remains incomplete.…

Biological Physics · Physics 2025-12-18 Sahil Islam , Mohd. Suhail Rizvi , Anupam Gupta

Unlike conventional two-dimensional (2D) semiconductor superlattices, moir\'{e} patterns in 2D materials are flexible and their electronic, magnetic, optical, and mechanical properties depend on their topography. Within a…

Mesoscale and Nanoscale Physics · Physics 2022-11-07 Alexandre Artaud , Nicolas Rougemaille , Sergio Vlaic , Vincent T. Renard , Nicolae Atodiresei , Johann Coraux

Active nematic models explain the topological defects and flow patterns observed in epithelial tissues, but the nature of active stress-whether it is extensile or contractile, a key parameter of the theory-is not well established…

Soft Condensed Matter · Physics 2025-11-11 Jan Rozman , Sumesh P. Thampi , Julia M. Yeomans

During epithelial wound healing, cell morphology near the healed wound and the healing rate vary strongly among different developmental stages even for a single species like \textit{Drosophila}. We develop deformable particle (DP) model…

Biological Physics · Physics 2024-02-23 Andrew T. Ton , Arthur K. MacKeith , Mark D. Shattuck , Corey S. O'Hern

We investigate morphologies of proliferating cellular tissue using a newly developed numerical simulation model for mechanical cell division. The model reproduces structures of simple multi-cellular organisms via simple rules for selective…

Soft Condensed Matter · Physics 2018-11-20 Pranav Madhikar , Jan Åström , Björn Baumeier , Mikko Karttunen

In this paper, we present a 2D numerical model developed to simulate the dynamics of soft, deformable particles. To accommodate significant particle deformations, the particle surface is represented as a narrow shell composed of mass points…

Computational Physics · Physics 2025-04-01 Yohann Trivino , Vincent Richefeu , Farhang Radjai , Komlanvi Lampoh , Jean-Yves Delenne

Space-saving design is a requirement that is encountered in biological systems and the development of modern technological devices alike. Many living organisms dynamically pack their polymer chains, filaments or membranes inside of…

Soft Condensed Matter · Physics 2014-07-18 Roman Vetter , Falk K. Wittel , Hans J. Herrmann

Biological processes, from morphogenesis to tumor invasion, spontaneously generate shear stresses inside living tissue. The mechanisms that govern the transmission of mechanical forces in epithelia and the collective response of the tissue…

Soft Condensed Matter · Physics 2022-11-29 Junxiang Huang , James O. Cochran , Suzanne M. Fielding , M. Cristina Marchetti , Dapeng Bi

Epithelial morphogenesis, a signature problem of tissue biology and tissue mechanics, continues to inspire biologists and physicists alike. Many treatments focus on tissue fluidization, apical/basal ratio changes, or mechanical…

Biological Physics · Physics 2023-07-26 Carlos M. Duque , Carl D. Modes

Morphogenesis involves complex shape changes of biological tissues. Yet, tissue shape changes depend on tissue rheology, which in turn arises from the interplay of large numbers of cells. Here, we link cell- and tissue-scale mechanics by…

Soft Condensed Matter · Physics 2026-02-27 Shalabh Kumar Anand , Matthias Merkel

Cell monolayers and epithelial tissues display slow dynamics during the liquid-glass transitions, a phenomenon with direct relevance to embryogenesis, tumor metastases, and wound healing. In active cells, persistent motion and cell…

Soft Condensed Matter · Physics 2024-01-25 Meng-Yuan Li , Yan-Wei Li

Over the last two decades, scientific literature has been blooming with various means of simulating epithelial cell colonies. Each of these simulations can be separated by their respective efficiency (expressed in terms of consumed…

Computational Physics · Physics 2022-12-20 Kevin Höllring , Ana-Sunčana Smith

Epithelial cell monolayers show remarkable displacement and velocity correlations over distances of ten or more cell sizes that are reminiscent of supercooled liquids and active nematics. We show that many observed features can be described…

Soft Condensed Matter · Physics 2020-04-22 Silke Henkes , Kaja Kostanjevec , J. Martin Collinson , Rastko Sknepnek , Eric Bertin

Surface tension governed by differential adhesion can drive fluid particle mixtures to sort into separate regions, i.e., demix. Does the same phenomenon occur in confluent biological tissues? We begin to answer this question for epithelial…