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Amoeboid cell migration is characterized by frequent changes of the direction of motion and resembles a persistent random walk on long time scales. Although it is well known that cell migration is typically driven by the actin cytoskeleton,…

Biological Physics · Physics 2021-06-09 Nicolas Ecker , Karsten Kruse

Biological cells sense and respond to mechanical forces, but how such a mechanosensing proccess takes place in a nonlinear inhomogeneous fibrous matrix remains unknown. We show that cells in a fibrous matrix induce deformation fields that…

Biological Physics · Physics 2015-06-10 Jacob Notbohm , Ayelet Lesman , Phoebus Rosakis , David A. Tirrell , Guruswami Ravichandran

The one-dimensional crawling movement of a cell is considered in this theoretical study. Our active gel model shows that for a cell with weakly mechanosensitive adhesion complexes, as myosin contractility increases, a cell starts to move at…

Soft Condensed Matter · Physics 2023-01-05 Jen-Yu Lo , Yuan-Heng Tseng , Hsuan-Yi Chen

Semiflexible polymers such as filamentous actin play a vital role in the mechanical behavior of cells, yet the basic properties of cross-linked F-actin networks remain poorly understood. To address this issue, we have performed numerical…

Soft Condensed Matter · Physics 2009-11-10 D. A. Head , A. J. Levine , F. C. MacKintosh

Actin networks, acting as an engine pushing against an external load, are fundamentally important to cell motility. A measure of the effectiveness of an engine is the velocity the engine is able to produce at a given force, the…

Subcellular Processes · Quantitative Biology 2011-11-30 Daniel B. Smith , Jian Liu

Cytoskeletal active nematics exhibit striking non-equilibrium dynamics that are powered by energy-consuming molecular motors. To gain insight into the structure and mechanics of these materials, we design programmable clusters in which…

Soft Condensed Matter · Physics 2021-09-15 Alexandra M. Tayar , Michael F. Hagan , Zvonimir Dogic

Contractile biopolymer networks, such as the actomyosin meshwork of animal cells, are ubiquitous in living organisms. The active gel theory, which provides the thermodynamic framework for these materials, has been mostly used in conjunction…

Soft Condensed Matter · Physics 2025-01-13 Antoine Jallon , Pierre Recho , Jocelyn Étienne

We propose a theory based on non-equilibrium thermodynamics to describe the mechanical behavior of an active polymer gel created by the inclusion of molecular motors in its solvent. When activated, these motors attach to the chains of the…

Soft Condensed Matter · Physics 2019-05-31 Mattia Bacca , Omar A. Saleh , Robert M. McMeeking

Contractility in animal cells is often generated by molecular motors such as myosin, which require polar substrates for their function. Motivated by recent experimental evidence of motor-independent contractility, we propose a robust…

Soft Condensed Matter · Physics 2022-05-04 Sihan Chen , Tomer Markovich , Fred C. MacKintosh

Mechanics has an important role during morphogenesis, both in the generation of forces driving cell shape changes and in determining the effective material properties of cells and tissues. Drosophila dorsal closure (DC) has emerged as a…

Motivated by recent experiments showing nonlinear elasticity of in vitro networks of the biopolymer actin cross-linked with filamin, we present an effective medium theory of flexibly cross-linked stiff polymer networks. We model such…

Soft Condensed Matter · Physics 2008-10-19 C. P. Broedersz , C. Storm , F. C. MacKintosh

The mechanical properties of the cell depend crucially on the tension of its cytoskeleton, a biopolymer network that is put under stress by active motor proteins. While the fibrous nature of the network is known to strongly affect the…

Soft Condensed Matter · Physics 2018-05-15 Pierre Ronceray , Chase P. Broedersz , Martin Lenz

Inside cells of living organisms, actin filaments and microtubules self-assemble and dissemble dynamically by incorporating actin or tubulin from the cell plasma or releasing it into their tips' surroundings. Such reaction-diffusion systems…

Biological Physics · Physics 2019-09-04 Johannes Pausch , Gunnar Pruessner

Optical tracking in vivo experiments reveal that diffusion of particles in biological cells is strongly enhanced in the presence of ATP and the experimental data for animal cells could previously be reproduced within a phenomenological…

Soft Condensed Matter · Physics 2017-03-22 Kento Yasuda , Ryuichi Okamoto , Shigeyuki Komura , Alexander S. Mikhailov

We present a novel flow instability that can arise in thin films of cytoskeletal fluids if the friction with the substrate on which the film lies is sufficiently strong. We consider a two dimensional, membrane-bound fragment containing…

Soft Condensed Matter · Physics 2009-11-13 A. C. Callan-Jones , J. -F. Joanny , J. Prost

The actin cortex is an active biopolymer network underneath the plasma membrane at the periphery of mammalian cells. It is a major regulator of cell shape through the generation of active cortical tension. In addition, the cortex…

Biological Physics · Physics 2023-05-15 Valentin Ruffine , Andreas Hartmann , Michael Schlierf , Elisabeth Fischer-Friedrich

The production of mechanical stresses in living organisms largely relies on localized, force-generating active units embedded in filamentous matrices. Numerical simulations of discrete fiber networks with fixed boundaries have shown that…

Biological Physics · Physics 2018-05-10 Pierre Ronceray , Chase Broedersz , Martin Lenz

We derive the constitutive equations of an active polar gel from a model for the dynamics of elastic molecules that link polar elements. Molecular binding kinetics induces the fluidization of the material, giving rise to Maxwell…

Biological Physics · Physics 2017-03-01 David Oriola , Ricard Alert , Jaume Casademunt

Regulation of contraction in striated muscle is controlled by a dual mechanism involving both thin filaments containing actin and thick filaments containing myosin. The thin filament is activated by calcium ions binding to troponin, leading…

The intracellular environment is a dynamic space filled with various organelles moving in all directions. Included in this diverse group of organelles are vesicles, which are involved in transport of molecular cargo throughout the cell.…

Biological Physics · Physics 2018-09-07 Danielle Posey , Paris Blaisdell-Pijuan , Samantha K. Knoll , Taher A. Saif , Wylie W. Ahmed