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Emergent order resulting from spontaneous symmetry breakings has been a central topic in statistical physics. Active matter systems composed of nonequilibrium elements exhibit a diverse range of fascinating phenomena beyond equilibrium…

Statistical Mechanics · Physics 2023-07-03 Daiki Nishiguchi

The cells and tissues that make up our body juggle contradictory mechanical demands. It is crucial for their survival to be able to withstand large mechanical loads, but it is equally crucial for them to produce forces and actively change…

Biological Physics · Physics 2019-03-22 Federica Burla , Yuval Mulla , Bart E. Vos , Anders Aufderhorst-Roberts , Gijsje H. Koenderink

The emergence of macroscopic order and patterns is a central paradigm in systems of (self-)propelled agents, and a key component in the structuring of many biological systems.The relationships between the ordering process and the underlying…

Soft Condensed Matter · Physics 2020-12-02 Jonas Denk , Erwin Frey

Active processes drive and guide biological dynamics across scales -- from subcellular cytoskeletal remodelling, through tissue development in embryogenesis, to population-level bacterial colonies expansion. In each of these, biological…

Light carries energy and momentum. It can therefore alter the motion of objects from atomic to astronomical scales. Being widely available, readily controllable and broadly biocompatible, light is also an ideal tool to propel microscopic…

Soft Condensed Matter · Physics 2025-07-22 Marcel Rey , Giovanni Volpe , Giorgio Volpe

Active matter systems such as eukaryotic cells and bacteria continuously transform chemical energy to motion. Hence living systems exert active stresses on the complex environments in which they reside. One recurring aspect of this…

Soft Condensed Matter · Physics 2020-02-27 Emmanuel Lance Christopher VI M. Plan , Julia Yeomans , Amin Doostmohammadi

Active matter consists of self-propelled elements exhibits fascinating collective motions ranging from biological to artificial systems. Among wide varieties of active matter systems, reconstituted bio-filaments moving on molecular motor…

Soft Condensed Matter · Physics 2020-03-25 Sakurako Tanida , Ken'ya Furuta , Kaori Nishikawa , Tetsuya Hiraiwa , Hiroaki Kojima , Kazuhiro Oiwa , Masaki Sano

The design of intelligent materials often draws parallels with the complex adaptive behaviors of biological organisms, where robust functionality stems from sophisticated hierarchical organization and emergent long-distance coordination…

Soft Condensed Matter · Physics 2025-11-13 Vladimir A. Baulin , Rudolf M. Füchslin , Achille Giacometti , Helmut Hauser , Marco Werner

Cooperative motion in biological microswimmers is crucial for their survival as it facilitates adhesion to surfaces, formation of hierarchical colonies, efficient motion, and enhanced access to nutrients. Synthetic microswimmers currently…

Soft Condensed Matter · Physics 2022-04-12 Stefania Ketzetzi , Melissa Rinaldin , Pim Dröge , Joost de Graaf , Daniela J. Kraft

We present a model of active particles interacting through a dynamic, heterogeneous environment, leading to emergent collective behaviors without direct agent-to-agent communication. Expanding the resource-dependent framework introduced in…

Biological Physics · Physics 2026-01-22 Gaston Briozzo , Gustavo J. Sibona , Fernando Peruani

Active systems are driven out of equilibrium by exchanging energy and momentum with their environment. This endows them with anomalous mechanical properties that we review in this colloquium for the case of dry scalar active matter, which…

Soft Condensed Matter · Physics 2025-01-20 Omer Granek , Yariv Kafri , Mehran Kardar , Sunghan Ro , Alexandre Solon , Julien Tailleur

Active matter systems may be characterised by the conversion of energy into active motion, e.g. the self-propulsion of microorganisms. Artificial active colloids form models which exhibit essential properties of more complex biological…

Soft Condensed Matter · Physics 2021-07-19 Abraham Mauleon-Amieva , Michael P. Allen , C. Patrick Royall

Active matter refers to a broad class of non-equilibrium systems where energy is continuously injected at the level of individual ``particles". These systems exhibit emergent collective behaviors that have no direct thermal-equilibrium…

Soft Condensed Matter · Physics 2026-02-10 Zihan Zheng , Cunyuan Jiang , Yangrui Chen , Matteo Baggioli , Jie Zhang

Biological molecules, like all active matter, use free energy to generate force and motion which drive them out of thermal equilibrium, and undergo inherent dynamic interconversion between metastable free energy states separated by levels…

Biological Physics · Physics 2025-08-28 Mark C Leake

Active matter has been intensely studied for its wealth of intriguing properties such as collective motion, motility-induced phase separation (MIPS), and giant fluctuations away from criticality. However, the precise connection of active…

Statistical Mechanics · Physics 2019-04-17 Juliane U. Klamser , Sebastian C. Kapfer , Werner Krauth

Active matter is one of the hottest topics in physics nowadays. As a prototype of living systems operating in viscous environments it has usually been modeled in terms of the overdamped dynamics. Recently, active matter in the underdamped…

Statistical Mechanics · Physics 2025-03-31 K. Białas , J. Spiechowicz

Active solids consist of elastically coupled out-of-equilibrium units performing work. They are central to autonomous processes, such as locomotion, self-oscillations and rectification, in biological systems,designer materials and robotics.…

Soft Condensed Matter · Physics 2025-01-17 P. Baconnier , D. Shohat , C. Hernandèz , C. Coulais , V. Démery , G. Düring , O. Dauchot

We derive a hydrodynamic description of metric-free active matter: starting from self-propelled particles aligning with neighbors defined by "topological" rules, not metric zones, -a situation advocated recently to be relevant for bird…

Soft Condensed Matter · Physics 2015-06-04 Anton Peshkov , Sandrine Ngo , Eric Bertin , Hugues Chaté , Francesco Ginelli

The interplay between active matter and its environment is central to understanding emergent behavior in biological and synthetic systems. Here, we show that coupling active nematic flows to small-amplitude deformations of a compliant…

Soft Condensed Matter · Physics 2026-01-19 Varun Venkatesh , Amin Doostmohammadi

One characteristic feature of soft matter systems is their strong response to external stimuli. As a consequence they are comparatively easily driven out of their ground state and out of equilibrium, which leads to many of their fascinating…

Soft Condensed Matter · Physics 2015-01-29 Andreas M. Menzel
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