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Inspired by dense contractile tissues, where cells are subject to periodic deformation, we formulate and study a generic hydrodynamic theory of pulsating active liquids. Combining mechanical and phenomenological arguments, we postulate that…

Soft Condensed Matter · Physics 2025-09-25 Tirthankar Banerjee , Thibault Desaleux , Jonas Ranft , Étienne Fodor

A common feature of biological self-organization is how active agents communicate with each other or their environment via chemical signaling. Such communications, mediated by self-generated chemical gradients, have consequences for both…

Soft Condensed Matter · Physics 2023-11-07 Babak Vajdi Hokmabad , Suropriya Saha , Jaime Agudo-Canalejo , Ramin Golestanian , Corinna C. Maass

An active chiral rotor is a spherical object that can generate chiral flows in a fluid by rotating about an axis. For example, if the flow around the upper hemisphere of the chiral rotor is in a clockwise direction, then the flow in the…

Soft Condensed Matter · Physics 2025-01-29 R. Maity , Snigdha Thakur , P. S. Burada

Hydrodynamical interactions of active micro-particles are pervasive in our planet's fluid environments. Hence, understanding the interactions of these self-propelled particles is essential for science and engineering. In this paper the…

Soft Condensed Matter · Physics 2022-12-28 Yonatan Ashenafi

Many fascinating properties of biological active matter crucially depend on the capacity of constituting entities to perform directed motion, e.g., molecular motors transporting vesicles inside cells or bacteria searching for food. While…

Statistical Mechanics · Physics 2023-06-21 Pietro Luigi Muzzeddu , Édgar Roldán , Andrea Gambassi , Abhinav Sharma

Self-propelled phoretic swimmers are generally studied in the laminar flow regime, where their low speed renders inertial effects negligible and trajectories highly predictable. This research tackles the challenge of propulsion in the…

Fluid Dynamics · Physics 2026-01-19 Alessandro Foradori , Paolo Bettotti

Ciliated microswimmers and flagellated bacteria alter their swimming trajectories to follow the direction of an applied electric field exhibiting electrotaxis. Both for matters of application and physical modelling, it is instructive to…

Soft Condensed Matter · Physics 2024-01-26 Carola M. Buness , Avi Rana , Corinna C. Maass , Ranabir Dey

Microorganisms can preferentially orient and move along gradients of a chemo-attractant (i.e., chemotax) while colonies of many microorganisms can collectively undergo complex dynamics in response to chemo-attractants that they themselves…

Biological Physics · Physics 2013-10-30 Enkeleida Lushi , Raymond E. Goldstein , Michael J. Shelley

We study the nonequilibrium stationary state of a one-dimensional inertial run-and-tumble particle (IRTP) trapped in a harmonic potential. We find that the presence of inertia leads to two distinct dynamical scenarios, namely, overdamped…

Statistical Mechanics · Physics 2025-05-21 Debraj Dutta , Anupam Kundu , Sanjib Sabhapandit , Urna Basu

Both natural and artificial small-scale swimmers may often self-propel in environments subject to complex geometrical constraints. While most past theoretical work on low-Reynolds number locomotion addressed idealised geometrical…

Fluid Dynamics · Physics 2017-11-16 Alexander Chamolly , Takuji Ishikawa , Eric Lauga

We develop the hydrodynamic theory of dry, polar ordered, active matter (``flocking") with autochemotaxis; i.e., self-propelled entities moving in the same direction, each emitting a substance which attracts the others (e.g., ants). We find…

Soft Condensed Matter · Physics 2024-01-22 Maxx Miller , John Toner

Synthetic microswimmers show great promise in biomedical applications such as drug delivery and microsurgery. Their locomotion, however, is subject to stringent constraints due to the dominance of viscous over inertial forces at low…

Fluid Dynamics · Physics 2020-07-15 Alan Cheng Hou Tsang , Pun Wai Tong , Shreyes Nallan , On Shun Pak

The biological fluids encountered by self-propelled cells display complex microstructures and rheology. We consider here the general problem of low-Reynolds number locomotion in a complex fluid. {Building on classical work on the transport…

Fluid Dynamics · Physics 2014-10-16 Eric Lauga

Ensembles of particles rotating in a two-dimensional fluid can exhibit chaotic dynamics yet develop signatures of hidden order. Such "rotors" are found in the natural world spanning vastly disparate length scales - from the rotor proteins…

Soft Condensed Matter · Physics 2022-03-09 Naomi Oppenheimer , David B. Stein , Matan Yah Ben Zion , Michael J. Shelley

Locomotion by shape changes (spermatozoon swimming, snake slithering, bird flapping) or gas expulsion (rocket firing) is assumed to require environmental interaction, due to conservation of momentum. As first noted in (Wisdom, 2003) and…

The dynamics of two-dimensional fluids confined within a random matrix of obstacles is investigated using both colloidal model experiments and molecular dynamics simulations. By varying fluid and matrix area fractions in the experiment, we…

Biological and artificial microswimmers often self-propel in external flows of vortical nature; relevant examples include algae in small-scale ocean eddies, spermatozoa in uterine peristaltic flows and bacteria in microfluidic devices. A…

Biological Physics · Physics 2022-11-14 Ivan Tanasijevic , Eric Lauga

Robots are becoming increasingly essential for traversing complex environments such as disaster areas, extraterrestrial terrains, and marine environments. Yet, their potential is often limited by mobility and adaptability constraints. In…

Robotics · Computer Science 2025-03-10 Nnamdi Chikere , John McElroy , Yasemin Ozkan-Aydin

Motivated by the observation of non-exponential run-time distributions of bacterial swimmers, we propose a minimal phenomenological model for taxis of active particles whose motion is controlled by an internal clock. The ticking of the…

Biological Physics · Physics 2020-07-23 Luis Gómez Nava , Robert Großmann , Marius Hintsche , Carsten Beta , Fernando Peruani

We exactly solve the nonequilibrium dynamics of a harmonically trapped self-propelled particle with anisotropic translational mobility in two dimensions, relevant to rodlike microswimmers and wheeled robots. The mean displacement and MSD…

Statistical Mechanics · Physics 2026-05-05 Amir Shee , P. S. Pal