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Zooplankton live in dynamic environments where turbulence may challenge their limited swimming abilities. How this interferes with fundamental behavioral processes remains elusive. We reconstruct simultaneously the trajectories of flow…

Fluid Dynamics · Physics 2020-12-03 François-Gaël Michalec , Itzhak Fouxon , Sami Souissi , Markus Holzner

We employ three numerical methods to explore the motion of low Reynolds number swimmers, modeling the hydrodynamic interactions by means of the Oseen tensor approximation, lattice Boltzmann simulations and multiparticle collision dynamics.…

Soft Condensed Matter · Physics 2007-05-23 David J. Earl , C. M. Pooley , J. F. Ryder , Irene Bredberg , J. M. Yeomans

The locomotion of swimming bacteria in simple Newtonian fluids can successfully be described within the framework of low Reynolds number hydrodynamics. The presence of polymers in biofluids generally increases the viscosity, which is…

Soft Condensed Matter · Physics 2019-08-12 Andreas Zöttl , Julia M. Yeomans

Many marine invertebrates have larval stages covered in linear arrays of beating cilia, which propel the animal while simultaneously entraining planktonic prey. These bands are strongly conserved across taxa spanning four major superphyla,…

Fluid Dynamics · Physics 2017-02-14 William Gilpin , Vivek N. Prakash , Manu Prakash

We describe a new numerical 3D relativistic hydrodynamical code, the results of validation tests, and a comparison with earlier, 2D studies. The 3D code has been used to study the deflection and precession of relativistic flows. We find…

Astrophysics · Physics 2009-11-07 Philip A. Hughes , Mark A. Miller , G. Comer Duncan

We study the fluid dynamics of two fish-like bodies with synchronised swimming patterns. Our studies are based on two-dimensional simulations of viscous incompressible flows. We distinguish between motion patterns that are externally…

Trajectories and conformations of uni- and multiflagellar bacteria are studied with a coarse-grained model of a cell comprised of elastic flagella connected to a cell body. The elasticities of both the hook protein (connecting cell body and…

Biological Physics · Physics 2018-11-07 Frank T. M. Nguyen , Michael D. Graham

Due to its great efficiency and maneuverability, undulatory swimming is the predominant form of locomotion in aquatic vertebrates. A myriad of animals of different species and sizes oscillate their bodies to propel themselves in aquatic…

Fluid Dynamics · Physics 2026-01-26 Jesús Sánchez-Rodríguez , Christophe Raufaste , Mederic Argentina

We present a theory of jellyfish swarm formation and exemplify it with simulations of active Brownian particles. The motivation for our analysis is the phenomenon of jellyfish blooms in the ocean and clustering of jellyfish in tank…

Adaptation and Self-Organizing Systems · Physics 2024-11-26 Erik Gengel , Zafrir Kuplik , Dror Angel , Eyal Heifetz

We study a linear inviscid model of a passively flexible swimmer with distributed flexibility, calculating its propulsive performance and optimal distributions of flexibility. The frequencies of actuation and mean stiffness ratios we…

Fluid Dynamics · Physics 2020-02-26 Daniel Floryan , Clarence W. Rowley

We experimentally study a scallop-like swimmer with reciprocally flapping wings in a nearly frictionless, cohesive granular medium consisting of hydrogel spheres. Significant locomotion is found when the swimmer's flapping frequency matches…

Soft Condensed Matter · Physics 2026-04-29 Hongyi Xiao , Jing Wang , Achim Sack , Ralf Stannarius , Thorsten Pöschel

A novel model of the wake of swimming fish is developed and incorporated into a dynamical model of a fish school to explore the effect of hydrodynamics on the emergent behavior in schooling fish. The model incorporates well-established…

Fluid Dynamics · Physics 2025-07-11 Ji Zhou , Jung-Hee Seo , Rajat Mittal

Fish swim with flexible fins that stand in stark contrast to the rigid propulsors of engineered vehicles. Using numerical simulations of the dynamics of flow-structure interaction, we have found that dorso-ventral deformation in flexible…

Fluid Dynamics · Physics 2026-04-16 Sushrut Kumar , Matthew J. McHenry , Jung-Hee Seo , Rajat Mittal

The move to high performance applications greatly increases the demand to produce large instantaneous fluid forces for high-speed maneuvering and improved power efficiency for sustained propulsion. Animals achieve remarkable feats of…

Fluid Dynamics · Physics 2016-09-22 G D Weymouth

Efficient locomotion is important for the evolution of complex life, yet the physical principles selecting specific swimming strokes often remain entangled with biological constraints. In viscous fluids, the scallop theorem constrains the…

Biological Physics · Physics 2026-03-11 Takahiro Kanazawa , Kenta Ishimoto , Kyogo Kawaguchi

Swimming, i.e., being able to advance in the absence of external forces by performing cyclic shape changes, is particularly demanding at low Reynolds numbers which is the regime of interest for micro-organisms and micro-robots. We focus on…

Optimization and Control · Mathematics 2012-10-04 François Alouges , Laetitia Giraldi

When a microorganism begins swimming from rest in a Newtonian fluid such as water, it rapidly attains its steady-state swimming speed since changes in the velocity field spread quickly when the Reynolds number is small. However, swimming…

Fluid Dynamics · Physics 2016-02-12 Madison S. Krieger , Marcelo A. Dias , Thomas R. Powers

Fish schools are ubiquitous in marine life. Although flow interactions are thought to be beneficial for schooling, their exact effects on the speed, energetics, and stability of the group remain elusive. Recent experiments suggest that flow…

Fluid Dynamics · Physics 2021-08-04 Sina Heydari , Eva Kanso

Reciprocal movement cannot be used for locomotion at low-Reynolds number in an infinite fluid or near a rigid surface. Here we show that this limitation is relaxed for a body performing reciprocal motions near a deformable interface. Using…

Soft Condensed Matter · Physics 2008-10-02 Renaud Trouilloud , Tony S. Yu , A. E. Hosoi , Eric Lauga

Both natural and artificial small-scale swimmers may often self-propel in environments subject to complex geometrical constraints. While most past theoretical work on low-Reynolds number locomotion addressed idealised geometrical…

Fluid Dynamics · Physics 2017-11-16 Alexander Chamolly , Takuji Ishikawa , Eric Lauga