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Marine organisms manipulate their surrounding flow through their swimming dynamics, which affects the transport of their own odor cues. We demonstrate by direct numerical simulations how a group of swimmers, moving at intermediate Reynolds…

Fluid Dynamics · Physics 2026-05-04 Martin James , Francesco Viola , Agnese Seminara

It has been a long-standing problem how schooling fish optimize their motion by exploiting the vortices shed by the others. A recent experimental study showed that a pair of fish reduce energy consumption by matching the phases of their…

Biological Physics · Physics 2023-12-05 Susumu Ito , Nariya Uchida

Researchers have long debated which spatial arrangements and swimming synchronizations are beneficial for the hydrodynamic performance of fish in schools. In our previous work (Seo and Mittal, Bioinsp. Biomim., Vol. 17, 066020, 2022), we…

Fluid Dynamics · Physics 2025-07-11 Ji Zhou , Jung-Hee Seo , Rajat Mittal

The stereotypical reaching motion of the octopus arm has drawn growing attention for its efficient control of a highly deformable body. Previous studies suggest that its characteristic bend propagation may share underlying principles with…

We examine the motion of rigid, ellipsoidal swimmers subjected to a steady vortex flow in two dimensions. Numerical simulations of swimmers in a spatially periodic array of vortices reveal a range of possible behaviors, including trapping…

Chaotic Dynamics · Physics 2020-08-11 S. A. Berman , K. A. Mitchell

We conduct experiments with flexible swimmers to address the impact of fluid viscoelasticity on their locomotion. The swimmers are composed of a magnetic head actuated in rotation by a frequency-controlled magnetic field and a flexible tail…

Fluid Dynamics · Physics 2013-03-19 Julian Espinosa-Garcia , Eric Lauga , Roberto Zenit

A single flexible filament can be actuated to escape from the scallop theorem and generate net propulsion at low Reynolds number. In this work, we study the dynamics of a simple boundary-driven multi-filament swimmer, a two-arm clamshell…

Fluid Dynamics · Physics 2022-03-07 Shiyuan Hu , Jun Zhang , Michael J. Shelley

Metachronal swimming, the sequential beating of limbs with a small phase lag, is observed in many organisms at various scales, but has been studied mostly in the limits of high or low Reynolds numbers. Motivated by the swimming of brine…

Fluid Dynamics · Physics 2021-08-03 Hong Nguyen , Daphne Klotsa

Most aquatic vertebrates swim by lateral flapping of their bodies and caudal fins. While much effort has been devoted to understanding the flapping kinematics and its influence on the swimming efficiency, little is known about the stability…

Fluid Dynamics · Physics 2015-06-15 Fangxu Jing , Eva Kanso

Over a sufficiently long period of time, or from an appropriate distance, the motion of many swimmers can appear smooth, with their trajectories appearing almost ballistic in nature and slowly varying in character. These long-time…

Soft Condensed Matter · Physics 2022-02-08 Benjamin J. Walker , Kenta Ishimoto , Eamonn A. Gaffney , Clément Moreau , Mohit P. Dalwadi

Fish schooling is often modeled with self-propelled particles subject to phenomenological behavioral rules. Although fish are known to sense and exploit flow features, these models usually neglect hydrodynamics. Here, we propose a novel…

Biological Physics · Physics 2018-05-16 Audrey Filella , François Nadal , Clément Sire , Eva Kanso , Christophe Eloy

Fish display remarkable swimming capabilities through the coordinated interaction of the body and caudal fin, yet the potential role of a passively pitching tail in enhancing hydrodynamic performance remains unresolved. In this work, we…

Fluid Dynamics · Physics 2025-11-06 Dev Pradeepkumar Nayak , Ali Tarokh , Muhammad Saif Ullah Khalid

Swimming microorganisms often self propel in fluids with complex rheology. While past theoretical work indicates that fluid viscoelasticity should hinder their locomotion, recent experiments on waving swimmers suggest a possible…

Biological Physics · Physics 2014-11-25 Emily E. Riley , Eric Lauga

Motivated by the aim of understanding the effect of media heterogeneity on the swimming dynamics of flagellated bacteria, we study the rotation and swimming of rigid helices in dilute suspensions experimentally and theoretically. We first…

Fluid Dynamics · Physics 2024-11-27 Albane Théry , Andres Zambrano , Eric Lauga , Roberto Zenit

We present a detailed numerical study of a microscopic artificial swimmer realized recently by Dreyfus et al. in experiments [R. Dreyfus et al., Nature 437, 862 (2005)]. It consists of an elastic filament composed of superparamagnetic…

Soft Condensed Matter · Physics 2008-05-21 Erik M. Gauger , Holger Stark

Fish perform various propulsive maneuvers while swimming by generating traveling waves along their bodies and producing thrust through tail strokes. Anguilliform swimmers spread motion along the body, while carangiform swimmers' motion is…

Fluid Dynamics · Physics 2025-12-04 Zahra Maleksabet , Maham Kamran , Ali Tarokh , Muhammad Saif Ullah Khalid

Active flexible filaments form the classical continuum framework for modelling the locomotion of spermatozoa and algae driven by the periodic oscillation of flagella. This framework also applies to the locomotion of various artificial…

Biological Physics · Physics 2025-10-14 John Severn , Eric Lauga

This study examines the impact of in-phase schooling on the hydrodynamic efficiency during linear acceleration in a simplified model using two undulating NACA0012 hydrofoils arranged in phalanx formation as a minimal representation of a…

Fluid Dynamics · Physics 2023-12-13 Zhonglu Lin , Dongfang Liang , Yu Zhang

Cell motility in viscous fluids is ubiquitous and affects many biological processes, including reproduction, infection, and the marine life ecosystem. Here we review the biophysical and mechanical principles of locomotion at the small…

Soft Condensed Matter · Physics 2009-09-16 Eric Lauga , Thomas R. Powers

Many swimming and flying animals are observed to cruise in a narrow range of Strouhal numbers, where the Strouhal number ${St = 2fA/U}$ is a dimensionless parameter that relates stroke frequency $f$, amplitude $A$, and forward speed $U$.…

Fluid Dynamics · Physics 2019-04-11 Daniel Floryan , Tyler Van Buren , Alexander J. Smits