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Related papers: Binary pusher-puller mixtures of active microswimm…

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Suspensions of rear- and front-actuated microswimmers immersed in a fluid, known respectively as ``pushers'' and ``pullers'', display qualitatively different collective behaviours: beyond a characteristic density, pusher suspensions exhibit…

Soft Condensed Matter · Physics 2020-07-08 Dóra Bárdfalvy , Shan Anjum , Cesare Nardini , Alexander Morozov , Joakim Stenhammar

Self-propelled micron-size particles suspended in a fluid, like bacteria or synthetic microswimmers, are strongly non-equilibrium systems where particle motility breaks the microscopic detailed balance, often resulting in large-scale…

Soft Condensed Matter · Physics 2024-03-15 Dóra Bárdfalvy , Viktor Škultéty , Cesare Nardini , Alexander Morozov , Joakim Stenhammar

Systems of motile microorganisms exhibit a multitude of collective phenomena, including motility-induced phase separation and turbulence. Sensing of the environment and adaptation of movement plays an essential role in the emergent…

Soft Condensed Matter · Physics 2024-06-26 Segun Goh , Elmar Westphal , Roland G. Winkler , Gerhard Gompper

Microswimmers typically operate in complex environments. In biological systems, often diverse species are simultaneously present and interact with each other. Here, we derive a (time-dependent) particle-scale statistical description, namely…

Soft Condensed Matter · Physics 2019-08-13 Christian Hoell , Hartmut Löwen , Andreas M. Menzel

We use numerical simulations to probe the dynamics of concentrated suspensions of spherical microswimmers interacting hydrodynamically. Previous work in the dilute limit predicted orientational instabilities of aligned suspensions for both…

Soft Condensed Matter · Physics 2013-02-12 Arthur A. Evans , Takuji Ishikawa , Takami Yamaguchi , Eric Lauga

Microorganisms living in microfluidic environments often form multi-species swarms, where they can leverage collective motions to achieve enhanced transport and spreading. Nevertheless, there is a general lack of physical understandings of…

Fluid Dynamics · Physics 2022-02-14 Bryce Palmer , Wen Yan , Tong Gao

Previous particle-based computer simulations have revealed a significantly more pronounced tendency of spontaneous global polar ordering in puller (contractile) microswimmer suspensions than in pusher (extensile) suspensions. We here…

Soft Condensed Matter · Physics 2018-10-23 Christian Hoell , Hartmut Löwen , Andreas M. Menzel

We explore the mechanism underlying the anomalous rheology of puller-type microswimmer suspensions through direct hydrodynamic simulations. Puller-type swimmers generate contractile flow fields along their swimming direction, leading to…

Soft Condensed Matter · Physics 2025-03-04 Haruki Hayano , Akira Furukawa

Microorganisms ofter move in confined, disordered environments, where hydrodynamic couplings can modify their transport behavior. Using extensive finite-element simulations, we investigate the dynamics of microswimmers -- modeled as…

Soft Condensed Matter · Physics 2026-03-24 Mirko Residori , Sebastian Aland , Christina Kurzthaler

Microswimmers in suspension exhibit collective swimming behaviour, forming various self-organised structures including ordered, aggregated, and turbulent-like structures. When mixed with passive particles phase-separation is known to occur,…

Fluid Dynamics · Physics 2026-03-26 Alexander Chamolly , Takuji Ishikawa

Few simulations exist for microswimmers near deformable interfaces. Here, we present numerical simulations of the hydrodynamic flows associated with a single microswimmer embedded in a binary fluid mixture. The two fluids demix, separated…

Soft Condensed Matter · Physics 2022-05-24 Chao Feng , John J. Molina , Matthew S. Turner , Ryoichi Yamamoto

Cooperation between micro-organisms give rise to novel phenomena like clustering, swarming in suspension. We study the collective behavior of the artificial swimmer called Taylor line at low Reynolds number using multi-particle collision…

Soft Condensed Matter · Physics 2018-02-14 Adyant Agrawal , Sujin B. Babu

Active matter has been the object of huge amount of research in recent years for its important fundamental and applicative properties. In this paper we investigate active suspensions of micro-swimmers through direct numerical simulation, so…

Soft Condensed Matter · Physics 2021-02-03 Sergio Chibbaro , Astrid Decoene , Sebastien Martin , Fabien Vergnet

A collection of microswimmers immersed in an incompressible fluid is characterised by strong interactions due to the long-range nature of the hydrodynamic fields generated by individual organisms. As a result, suspensions of rear-actuated…

Soft Condensed Matter · Physics 2023-12-19 Viktor Škultéty , Dóra Bárdfalvy , Joakim Stenhammar , Cesare Nardini , Alexander Morozov

The habitat of microorganisms is typically complex and viscoelastic. A natural question arises: Do polymers in a suspension of active swimmers enhance chaotic motion or promote orientational order? We address this issue by performing…

Soft Condensed Matter · Physics 2025-12-23 Yuan Zhou , Qingzhi Zou , Ignacio Pagonabarraga , Kaihuan Zhang , Kai Qi

An artificial microswimmer drifts in response to spatio-temporal modulations of an activating suspension medium. We consider two competing mechanisms capable of influencing its tactic response: angular fluctuations, which help it explore…

Soft Condensed Matter · Physics 2017-02-15 Alexander Geiseler , Peter Hänggi , Fabio Marchesoni

Suspensions of unicellular microswimmers such as flagellated bacteria or motile algae exhibit spontaneous density heterogeneities at large enough concentrations. Based on the relative location of the biological actuation appendages i.e.…

Soft Condensed Matter · Physics 2017-11-27 Fabian Jan Schwarzendahl , Marco G. Mazza

Self-propulsion of cellular microswimmers generates flow signatures, commonly classified as pusher- and puller-type, which characterize hydrodynamic interactions with other cells or boundaries. Using experimentally measured beat patterns,…

Cell Behavior · Quantitative Biology 2016-01-20 Gary S. Klindt , Benjamin M. Friedrich

Many microswimmers are inherently chiral, and this chirality can introduce fascinating behaviors in a collection of microswimmers. The dynamics become even more intriguing when two types of microswimmers with distinct chirality are mixed.…

Soft Condensed Matter · Physics 2025-09-03 Divya Kushwaha , Shradha Mishra

The ability of microswimmers to deploy optimal propulsion strategies is of paramount importance for their locomotory performance and survival at low Reynolds numbers. Although for perfectly spherical swimmers minimum dissipation requires a…

Fluid Dynamics · Physics 2021-07-13 Abdallah Daddi-Moussa-Ider , Babak Nasouri , Andrej Vilfan , Ramin Golestanian
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