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We study time-periodic forcing of spatially-extended patterns near a Turing-Hopf bifurcation point. A symmetry-based normal form analysis yields several predictions, including that (i) weak forcing near the intrinsic Hopf frequency enhances…

Pattern Formation and Solitons · Physics 2015-05-13 C. M. Topaz , Anne J. Catlla

Turing patterns are a cornerstone of biological self-organization, yet their emergence typically requires finely tuned parameters occupying narrow regions of high-dimensional space. This poses a fundamental challenge: how can evolving…

Cell Behavior · Quantitative Biology 2026-05-18 Seyoon Kim , Antonio Matas-Gil , Robert G. Endres

We consider the pattern formation problem in coupled identical systems after the global synchronized state becomes unstable. Based on analytical results relating the coupling strengths and the instability of each spatial mode (pattern) we…

Pattern Formation and Solitons · Physics 2009-11-10 Govindan Rangarajan , Yonghong Chen , Mingzhou Ding

The Brusselator is a generic reaction-diffusion model for a tri-molecular chemical reaction. We consider the case when the input and output reactions are slow. In this limit, we show the existence of $K$-periodic, spatially bi-stable…

Pattern Formation and Solitons · Physics 2009-11-11 Theodore Kolokolnikov , Thomas Erneux , Juncheng Wei

A class of systems is considered, where immobile species associated to distinct patches, the nodes of a network, interact both locally and at a long-range, as specified by an (interaction) adjacency matrix. Non local interactions are…

Pattern Formation and Solitons · Physics 2020-06-30 Giulia Cencetti , Federico Battiston , Timoteo Carletti , Duccio Fanelli

The aim of this paper is to contribute to the understanding of the pattern formation phenomenon in reaction-diffusion equations coupled with ordinary differential equations. Such systems of equations arise, for example, from modeling of…

Analysis of PDEs · Mathematics 2016-07-15 Anna Marciniak-Czochra , Grzegorz Karch , Kanako Suzuki

We study Turing pattern formation in a system undergoing radial growth in two dimensions. The Lengyel-Epstein two variable model is implemented in COMSOL Multiphysics and solved on domains growing at different speeds while sweeping other…

Pattern Formation and Solitons · Physics 2019-12-10 Noah H. Somberg , Christopher Konow , Irving R. Epstein , Milos Dolnik

We derive a necessary and sufficient condition for Turing instabilities to occur in two-component systems of reaction-diffusion equations with Neumann boundary conditions. We apply this condition to reaction-diffusion systems built from…

Mathematical Physics · Physics 2007-05-23 Rui Dilao

The effect of multiplicative noise to the Turing instability of the Brusselator system is investigated. We show that when the noise acts on both of the concentrations with the same intensities, then the Turing instability is suppressed…

Analysis of PDEs · Mathematics 2025-03-24 Qasim Khan , Anthony Suen , Bao Quoc Tang

On the one hand, checking specific termination proofs by hand, say using a particular collection of matrix interpretations, can be an arduous and error-prone task. On the other hand, automation of such checks would save time and help to…

Logic in Computer Science · Computer Science 2018-06-14 Jonas Schöpf , Christian Sternagel

Accurate and robust spatial orders are ubiquitous in living systems. In 1952, Alan Turing proposed an elegant mechanism for pattern formation based on spontaneous breaking of the spatial translational symmetry in the underlying…

Statistical Mechanics · Physics 2022-06-07 Dongliang Zhang , Chenghao Zhang , Qi Ouyang , Yuhai Tu

Turing's theory of pattern formation is a universal model for self-organization, applicable to many systems in physics, chemistry and biology. Essential properties of a Turing system, such as the conditions for the existence of patterns and…

Adaptation and Self-Organizing Systems · Physics 2018-08-02 Xavier Diego , Luciano Marcon , Patrick Müller , James Sharpe

Mutualisms are key for structuring ecological communities, but they are sensitive to environmental change and fluctuations in population size. Consequently, how mutualisms achieve stability remains an open question in ecological theory.…

Populations and Evolution · Quantitative Biology 2026-05-08 Matheus Bongestab , David Pinto-Ramos , Ricardo Martinez-Garcia

Symmetry-breaking instabilities play an important role in understanding the mechanisms underlying the diversity of patterns observed in nature, such as in Turing's reaction--diffusion theory, which connects cellular signalling and transport…

Pattern Formation and Solitons · Physics 2023-12-25 Andrew L. Krause , Eamonn A. Gaffney , Thomas Jun Jewell , Václav Klika , Benjamin J. Walker

Turing theory of pattern formation is among the most popular theoretical means to account for the variety of spatio-temporal structures observed in Nature and, for this reason, finds applications in many different fields. While Turing…

Pattern Formation and Solitons · Physics 2025-10-22 Riccardo Muolo , Luca Gallo , Vito Latora , Mattia Frasca , Timoteo Carletti

Phase separation in passive systems leads to uncontrolled droplet growth, limiting structural control in soft materials and cells. We identify a generic mechanism to arrest coarsening based on chemical interconversion between molecular…

Soft Condensed Matter · Physics 2026-03-19 Florian Raßhofer , Erwin Frey

Traditional linear stability analysis based on matrix diagonalization is a computationally intensive $O(n^3)$ process for $n$-dimensional systems of differential equations, posing substantial limitations for the exploration of Turing…

Quantitative Methods · Quantitative Biology 2023-09-12 Roozbeh H. Pazuki , Robert G. Endres

A characteristic feature of topological systems is the presence of robust gapless edge states. In this work the effect of time-dependent perturbations on the edge states is considered. Specifically we consider perturbations that can be…

Superconductivity · Physics 2017-01-16 P. D. Sacramento

As proposed by Alan Turing in 1952 as a ubiquitous mechanism for nonequilibrium pattern formation, diffusional effects may destabilize uniform distributions of reacting chemical species and lead to both spatially and temporally…

Pattern Formation and Solitons · Physics 2013-10-28 Shigefumi Hata , Hiroya Nakao , Alexander S. Mikhailov

We provide a constructive method designed in order to control the stability of a given periodic orbit of a general completely integrable system. The method consists of a specific type of perturbation, such that the resulting perturbed…

Dynamical Systems · Mathematics 2015-03-04 Razvan M. Tudoran