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Eukaryotic swimming cells such as spermatozoa, algae or protozoa use flagella or cilia to move in viscous fluids. The motion of their flexible appendages in the surrounding fluid induces propulsive forces that balance with the viscous drag…

Soft Condensed Matter · Physics 2023-10-06 Maciej Lisicki , Marcos F. Velho Rodrigues , Eric Lauga

A major puzzle in biology is how mammalian sperm determine and maintain the correct swimming direction during the various phases of the sexual reproduction process. Whilst chemotaxis is assumed to dominate in the immediate vicinity of the…

Biological Physics · Physics 2014-05-27 Vasily Kantsler , Jörn Dunkel , Martyn Blayney , Raymond E. Goldstein

We study the coupled hydrodynamics between a motile slender microswimmer and a deformable interface that separates two Newtonian fluid regions. From the disturbance field generated by the swimming motion, we quantitatively characterize the…

Soft Condensed Matter · Physics 2022-12-22 Sankalp Nambiar , John S Wettlaufer

Understanding the collective actuation of microscopic structures driven by external fields can lead to the development of next-generation autonomous machines. With this goal in mind, we investigated light-induced collective motion of…

Self-propelled phoretic colloids have recently emerged as a promising avenue for the design of artificial swimmers. These swimmers combine purely phoretic interactions with intricate hydrodynamics which critically depend on the swimmer…

Soft Condensed Matter · Physics 2018-01-17 Martin Wagner , Marisol Ripoll

Biological microswimmers often encounter deformable boundaries in physiological conditions; for instance, the viscoelastic walls of reproductive tract during migration of spermatozoa, or host tissue during early bacterial biofilm formation.…

Soft Condensed Matter · Physics 2025-08-07 Smita S. Sontakke , Aneesha Kajampady , Mohd Suhail Rizvi , Ranabir Dey

We analyze a minimal model for a rigid spherical microswimmer and explore the consequences of its extended surface on the interplay between its self-propulsion and flow properties. The model is the first order representation of…

Soft Condensed Matter · Physics 2017-12-06 Tapan Chandra Adhyapak , Sara Jabbari-Farouji

The dynamic interactions between pairs of swimming microorganisms underpin the collective behaviour of larger suspensions, but accurately calculating pairwise collisions has typically required the use of numerical simulations in which…

Soft Condensed Matter · Physics 2021-11-30 C. Darveniza , T. Ishikawa , T. J. Pedley , D. R. Brumley

Several micro-organisms, such as bacteria, algae, or spermatozoa, use flagella or cilia to swim in a fluid, while many other micro-organisms instead use ample shape deformation, described as amoeboid, to propel themselves by either crawling…

Biological Physics · Physics 2016-09-19 Hao Wu , A. Farutin , W. -F. Hu , M. Thiébaud , S. Rafaï , P. Peyla , M. -C. Lai , C. Misbah

We study the three-dimensional dynamics of a spherical microswimmer in cylindrical Poiseuille flow which can be mapped onto a Hamiltonian system. Swinging and tumbling trajectories are identified. In 2D they are equivalent to oscillating…

Soft Condensed Matter · Physics 2012-06-18 Andreas Zöttl , Holger Stark

We present new constrained and free-swimming experiments and simulations of a pair of pitching hydrofoils interacting in a minimal school. The hydrofoils have an out-of-phase synchronization and they are varied through in-line, staggered,…

Fluid Dynamics · Physics 2023-08-01 Pedro Costa Ormonde , Melike Kurt , Amin Mivehchi , Keith W. Moored

When groups of inertial swimmers move together, hydrodynamic interactions play a key role in shaping their collective dynamics, including the cohesion of the group. To explore how these interactions influence group cohesion, we develop a…

Fluid Dynamics · Physics 2025-05-23 Mohamed Niged Mabrouk , Daniel Floryan

Swimming cells and microorganisms are as diverse in their collective dynamics as they are in their individual shapes and propulsion mechanisms. Even for sperm cells, which have a stereotyped shape consisting of a cell body connected to a…

Fluid Dynamics · Physics 2018-06-25 S. F. Schoeller , E. E. Keaveny

Synchronization is one of the emerging collective phenomena in interacting particle systems. Its ubiquitous presence in nature, science, and technology has fascinated the scientific community over the decades. Moreover, a great deal of…

Adaptation and Self-Organizing Systems · Physics 2023-12-18 Suvam Pal , Gourab Kumar Sar , Dibakar Ghosh , Arnab Pal

The hydrodynamic flow field generated by self-propelled active particles and swimming microorganisms is strongly altered by the presence of nearby boundaries in a viscous flow. Using a simple model three-linked sphere swimmer, we show that…

Fluid Dynamics · Physics 2018-04-18 Abdallah Daddi-Moussa-Ider , Maciej Lisicki , Christian Hoell , Hartmut Löwen

The mechanism of hydrodynamics-induced pairing of soft particles, namely closed bilayer membranes (vesicles, a model system for red blood cells) and drops, is studied numerically with a special attention paid to the role of the confinement…

Few simulations exist for microswimmers near deformable interfaces. Here, we present numerical simulations of the hydrodynamic flows associated with a single microswimmer embedded in a binary fluid mixture. The two fluids demix, separated…

Soft Condensed Matter · Physics 2022-05-24 Chao Feng , John J. Molina , Matthew S. Turner , Ryoichi Yamamoto

To explore and react to their environment, living micro-swimmers have developed sophisticated strategies for locomotion - in particular, motility with multiple gaits. To understand the physical principles associated with such a behavioural…

Colloidal particles moving in a fluid interact via the induced velocity field. The collective dynamic state for a class of actively forced colloids, driven by harmonic potentials via a rule that couples forces to configurations, to perform…

Soft Condensed Matter · Physics 2011-10-11 Loïc Damet , Giovanni M. Cicuta , Jurij Kotar , Marco Cosentino Lagomarsino , Pietro Cicuta

Three-dimensional simulations with fully resolved hydrodynamics are performed to study the collective motion of model swimmers in confinement. We show that certain swimming mechanisms can lead to traveling wave-like collective motion even…

Soft Condensed Matter · Physics 2016-04-20 Norihiro Oyama , John Jairo Molina , Ryoichi Yamamoto