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Motivated by recent advances in vesicle engineering, we consider theoretically the locomotion of shape-changing bilayer vesicles at low Reynolds number. By modulating their volume and membrane composition, the vesicles can be made to change…

Biological Physics · Physics 2010-04-09 Arthur A. Evans , Saverio E. Spagnolie , Eric Lauga

Microorganisms ofter move in confined, disordered environments, where hydrodynamic couplings can modify their transport behavior. Using extensive finite-element simulations, we investigate the dynamics of microswimmers -- modeled as…

Soft Condensed Matter · Physics 2026-03-24 Mirko Residori , Sebastian Aland , Christina Kurzthaler

A simplified, pseudoplastic rheology characterized by constant viscosity plateaus above and below a transition strain rate is applied to axisymmetric, gravitationally driven spreading of a thin fluid film with constant volume flux source in…

Fluid Dynamics · Physics 2023-06-01 Chris Reese

We present a scheme of self-propelling liquid droplets which closely mimics the locomotion of some protozoal organisms, so-called squirmers. In contrast to other schemes proposed earlier, locomotion paths are not self-avoiding, since the…

Soft Condensed Matter · Physics 2011-03-29 Shashi Thutupalli , Ralf Seemann , Stephan Herminghaus

Hydrodynamic interaction strongly influences the collective behavior of the microswimmers. With this work, we study the behavior of two hydrodynamically interacting self-propelled chiral swimmers in the low Reynolds number regime,…

Soft Condensed Matter · Physics 2022-11-28 Ruma Maity , P. S. Burada

Swimming cells and microorganisms must often move though complex fluids that contain an immersed microstructure such as polymer molecules, or filaments. In many important biological processes, such as mammalian reproduction and bacterial…

Fluid Dynamics · Physics 2018-08-06 Arshad Kamal , Eric E Keaveny

A well-developed method to induce mixing on microscopic scales is to exploit flows generated by steady streaming. Steady streaming is a classical fluid dynamics phenomenon whereby a time-periodic forcing in the bulk or along a boundary is…

Fluid Dynamics · Physics 2015-12-02 Tamsin A. Spelman , Eric Lauga

Microswimmers are encountered in a wide variety of biophysical settings. When interacting with flow fields, they show interesting dynamical features such as trapping, clustering, and preferential orientation. One important step towards the…

Fluid Dynamics · Physics 2020-10-02 José-Agustín Arguedas-Leiva , Michael Wilczek

The locomotion of microorganisms and spermatozoa in complex viscoelastic fluids is of critical importance in many biological processes such as fertilization, infection, and biofilm formation. Depending on their propulsion mechanisms,…

Soft Condensed Matter · Physics 2021-09-14 Gaojin Li , Eric Lauga , Arezoo M. Ardekani

We present a lattice Boltzmann study of the hydrodynamics of a fully resolved squirmer, radius R, confined in a slab of fluid between two no-slip walls. We show that the coupling between hydrodynamics and short-range repulsive interactions…

Soft Condensed Matter · Physics 2016-03-30 Juho S. Lintuvuori , Aidan T. Brown , Kevin Stratford , Davide Marenduzzo

The incessant swimming motion of microbes in dense suspensions can give rise to striking collective motions and coherent structures. However, theoretical investigations of these structures typically utilize either computationally demanding…

Fluid Dynamics · Physics 2019-05-29 Douglas R. Brumley , Timothy J. Pedley

The dynamic interactions between pairs of swimming microorganisms underpin the collective behaviour of larger suspensions, but accurately calculating pairwise collisions has typically required the use of numerical simulations in which…

Soft Condensed Matter · Physics 2021-11-30 C. Darveniza , T. Ishikawa , T. J. Pedley , D. R. Brumley

The macroscopic properties of polymeric fluids are inherited from the material properties of the fibers embedded in the solvent. The behavior of such passive fibers in flow has been of interest in a wide range of systems, including cellular…

Soft Condensed Matter · Physics 2019-06-20 John LaGrone , Ricardo Cortez , Wen Yan , Lisa Fauci

We propose a hydrodynamic model for a spheroidal microswimmer with two tangential surface velocity modes. This model is analytically solvable and reduces to Lighthill's and Blake's spherical squirmer model in the limit of equal major and…

Biological Physics · Physics 2024-06-03 Mario Theers , Elmar Westphal , Gerhard Gompper , Roland G. Winkler

A small spheroid settling in a quiescent fluid experiences an inertial torque that aligns it so that it settles with its broad side first. Here we show that an active particle experiences such a torque too, as it settles in a fluid at rest.…

Fluid Dynamics · Physics 2023-12-21 F. Candelier , J. Qiu , L. Zhao , G. Voth , B. Mehlig

Autonomous and driven transport in chiral active fluids have been shown to exhibit features that cannot be accommodated within the classical formulation of fluid mechanics, due to the role of odd viscosity. We generalize the theory of…

Soft Condensed Matter · Physics 2026-04-24 Michalis Chatzittofi , Yuto Hosaka , Andrej Vilfan , Ramin Golestanian

In this work, we analyze the motion of an active particle, modeled as a spherical squirmer, in linearly varying viscosity fields. In general, the presence of a particle will disturb a background viscosity field and the disturbance generated…

Fluid Dynamics · Physics 2021-07-09 Vaseem A. Shaik , Gwynn J. Elfring

We run numerical simulations of strongly confined suspensions of model spherical swimmers called "squirmers". Because of the confinement, the Stokeslet dipoles generated by the particles are quickly screened and the far-field flow is…

Fluid Dynamics · Physics 2016-06-22 Jean-Baptiste Delfau , John Molina , Masaki Sano

We investigate how shear influences the dynamics of active particles in dilute to concentrated suspensions. Using apolar active suspensions of squirmers as model systems, we show how their long-time diffusive dynamics can surprisingly slow…

Soft Condensed Matter · Physics 2025-11-14 Zhouyang Ge , John F. Brady , Gwynn J. Elfring

Microswimmers display an intriguing ability to navigate through fluids with spatially varying viscosity, a behavior known as viscotaxis, which plays a crucial role in guiding their motion. In this study, we reveal that the orientation…

Fluid Dynamics · Physics 2025-06-25 Takuya Kobayashi , Ryoichi Yamamoto