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Biological evolution can be conceptualized as a search process in the space of gene sequences guided by the fitness landscape, a mapping that assigns a measure of reproductive value to each genotype. Here we discuss probabilistic models of…

Populations and Evolution · Quantitative Biology 2024-04-10 Joachim Krug , Daniel Oros

A fitness landscape is a mapping from a space of discrete genotypes to the real numbers. A path in a fitness landscape is a sequence of genotypes connected by single mutational steps. Such a path is said to be accessible if the fitness…

Populations and Evolution · Quantitative Biology 2023-02-21 Benjamin Schmiegelt , Joachim Krug

Functional effects of different mutations are known to combine to the total effect in highly nontrivial ways. For the trait under evolutionary selection (`fitness'), measured values over all possible combinations of a set of mutations yield…

Populations and Evolution · Quantitative Biology 2015-05-27 Jasper Franke , Alexander Klözer , J. Arjan G. M. de Visser , Joachim Krug

These notes introduce probabilistic landscape models defined on high-dimensional discrete sequence spaces. The models are motivated primarily by fitness landscapes in evolutionary biology, but links to statistical physics and computer…

Populations and Evolution · Quantitative Biology 2025-12-24 Sakshi Pahujani , Joachim Krug

The adaptive evolution of a population under the influence of mutation and selection is strongly influenced by the structure of the underlying fitness landscape, which encodes the interactions between mutations at different genetic loci.…

Populations and Evolution · Quantitative Biology 2015-06-04 Jasper Franke , Joachim Krug

All possible phenotypes are not equally accessible to evolving populations. In fact, only phenotypes of large size, i.e. those resulting from many different genotypes, are found in populations of sequences, presumably because they are…

Populations and Evolution · Quantitative Biology 2023-08-16 Pablo Catalán , Juan Antonio García-Martín , Jacobo Aguirre , José A. Cuesta , Susanna Manrubia

Fitness landscapes are mappings between genotypes, phenotypes, and fitness that shape evolution. In recent years, empirical work and theoretical models have greatly advanced our understanding of how populations navigate rugged fitness…

Populations and Evolution · Quantitative Biology 2026-04-21 Malvika Srivastava , Claudia Bank , Joachim Krug , Suman G. Das

Accessibility percolation is a new type of percolation problem inspired by evolutionary biology. To each vertex of a graph a random number is assigned and a path through the graph is called accessible if all numbers along the path are in…

Statistical Mechanics · Physics 2013-04-04 Stefan Nowak , Joachim Krug

We introduce a new model of evolution on a fitness landscape possessing a tunable degree of neutrality. The model allows us to study the general properties of molecular species undergoing neutral evolution. We find that a number of…

adap-org · Physics 2007-05-23 M. E. J. Newman , Robin Engelhardt

A fitness landscape is a mapping from the space of genetic sequences, which is modeled here as a binary hypercube of dimension $L$, to the real numbers. We consider random models of fitness landscapes, where fitness values are assigned…

Populations and Evolution · Quantitative Biology 2015-06-16 Benjamin Schmiegelt , Joachim Krug

We study how correlations in the random fitness assignment may affect the structure of fitness landscapes. We consider three classes of fitness models. The first is a continuous phenotype space in which individuals are characterized by a…

Populations and Evolution · Quantitative Biology 2007-05-23 Janko Gravner , Damien Pitman , Sergey Gavrilets

Accessibility percolation is a new type of percolation problem inspired by evolutionary biology: a random number, called its fitness, is assigned to each vertex of a graph, then a path in the graph is accessible if fitnesses are strictly…

Probability · Mathematics 2023-03-01 Frank Duque , Daniel Ramirez-Gomez , Alejandro Roldán-Correa , Leon A. Valencia

Consider a graph in which each site is endowed with a value called \emph{fitness}. A path in the graph is said to be "open" or "accessible" if the fitness values along that path is strictly increasing. We say that there is accessibility…

Probability · Mathematics 2014-01-28 Julien Berestycki , Éric Brunet , Zhan Shi

The tempo and mode of an adaptive process is strongly determined by the structure of the fitness landscape that underlies it. In order to be able to predict evolutionary outcomes (even on the short term), we must know more about the nature…

Populations and Evolution · Quantitative Biology 2013-02-28 Bjørn Østman , Christoph Adami

The idea of adaptive walks on fitness landscapes as a means of studying evolutionary processes on large time scales is extended to fitness landscapes that are slowly changing over time. The influence of ruggedness and of the amount of…

Biological Physics · Physics 2009-10-31 Claus O. Wilke , Thomas Martinetz

Random walks on multidimensional nonlinear landscapes are of interest in many areas of science and engineering. In particular, properties of adaptive trajectories on fitness landscapes determine population fates and thus play a central role…

Populations and Evolution · Quantitative Biology 2014-10-08 Michael Manhart , Alexandre V. Morozov

We consider the evolutionary trajectories traced out by an infinite population undergoing mutation-selection dynamics in static, uncorrelated random fitness landscapes. Starting from the population that consists of a single genotype, the…

Populations and Evolution · Quantitative Biology 2009-11-11 Kavita Jain , Joachim Krug

Evolutionary adaptation is the process that increases the fit of a population to the fitness landscape it inhabits. As a consequence, evolutionary dynamics is shaped, constrained, and channeled, by that fitness landscape. Much work has been…

Populations and Evolution · Quantitative Biology 2010-12-17 Bjørn Østman , Arend Hintze , Christoph Adami

Inspired by biological evolution, we consider the following so-called accessibility percolation problem: The vertices of the unoriented $n$-dimensional binary hypercube are assigned independent $U(0, 1)$ weights, referred to as fitnesses. A…

Probability · Mathematics 2015-01-12 Anders Martinsson

We investigate an simple evolutionary game of sequences and demonstrate on this example the structure of fitness landscapes in discrete problems. We show the smoothing action of the genotype-phenotype mapping which still makes it feasible…

adap-org · Physics 2008-02-03 Torsten Asselmeyer , Werner Ebeling , Helge Rose
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