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Microswimmers, especially in theoretical treatments, are generally taken to be completely inertia-free, since inertial effects on their motion are typically small and assuming their absence simplifies the problem considerably. Yet in nature…

Soft Condensed Matter · Physics 2016-11-08 Jayant Pande , Kristina Pickl , Oleg Trosman , Ulrich Rüde , Ana-Sunčana Smith

A striking feature of the collective behavior of spherical microswimmers is that for sufficiently strong self-propulsion they phase-separate into a dense cluster coexisting with a low-density dis- ordered surrounding. Extending our previous…

Soft Condensed Matter · Physics 2016-10-31 Johannes Blaschke , Maurice Maurer , Karthik Menon , Andreas Zöttl , Holger Stark

We study the fluid dynamics of two fish-like bodies with synchronised swimming patterns. Our studies are based on two-dimensional simulations of viscous incompressible flows. We distinguish between motion patterns that are externally…

The dynamics and motion of multi-ciliated microswimmers with a spherical body and a small number N (with 5 < N < 60) of cilia with length comparable to the body radius, is investigated by mesoscale hydrodynamics simulations. A metachronal…

Soft Condensed Matter · Physics 2024-06-03 Sebastian Rode , Jens Elgeti , Gerhard Gompper

We investigate theoretically the collective dynamics of a suspension of low Reynolds number swimmers that are confined to two dimensions by a thin fluid film. Our model swimmer is characterized by internal degrees of freedom which locally…

Biological Physics · Physics 2015-05-19 M. Leoni , T. B. Liverpool

We analyse the effect of confining rigid and elastic boundaries on the motility of a model dipolar microswimmer. Flexible boundaries are deformed by the velocity field of the swimmer in such a way that the motility of both extensile and…

Soft Condensed Matter · Physics 2015-06-15 Rodrigo Ledesma-Aguilar , Julia M. Yeomans

Swimming micro-organisms such as bacteria or spermatozoa are typically found in dense suspensions, and exhibit collective modes of locomotion qualitatively different from that displayed by isolated cells. In the dilute limit where…

Biological Physics · Physics 2010-05-02 Sebastien Michelin , Eric Lauga

Actuating periodically an elastic filament in a viscous liquid generally breaks the constraints of Purcell's scallop theorem, resulting in the generation of a net propulsive force. This observation suggests a method to design simple…

Soft Condensed Matter · Physics 2009-09-29 Eric Lauga

We show using theory and experiments that a small particle moving along an elastic membrane through a viscous fluid is repelled from the membrane due to hydro-elastic forces. The viscous stress field produces an elastic disturbance leading…

Soft Condensed Matter · Physics 2018-02-23 Bhargav Rallabandi , Naomi Oppenheimer , Matan Yah Ben Zion , Howard A. Stone

Using a fluid-particle dynamics approach, we numerically study the effects of hydrodynamic interactions on the collective dynamics of active suspensions within a simple model for bacterial motility: each microorganism is modeled as a…

Soft Condensed Matter · Physics 2015-06-22 Akira Furukawa , Davide Marenduzzo , Michael E Cates

Efficient locomotion is important for the evolution of complex life, yet the physical principles selecting specific swimming strokes often remain entangled with biological constraints. In viscous fluids, the scallop theorem constrains the…

Biological Physics · Physics 2026-03-11 Takahiro Kanazawa , Kenta Ishimoto , Kyogo Kawaguchi

We study the properties and symmetries governing the hydrodynamic interaction between two identical, arbitrarily shaped objects, driven through a viscous fluid. We treat analytically the leading (dipolar) terms of the pair-mobility matrix,…

Fluid Dynamics · Physics 2016-05-03 Tomer Goldfriend , Haim Diamant , Thomas A. Witten

The journey of mammalian spermatozoa in nature is well-known to be reliant on their individual motility. Often swimming in crowded microenvironments, the progress of any single swimmer is likely dependent on their interactions with other…

Fluid Dynamics · Physics 2019-09-11 Benjamin J. Walker , Kenta Ishimoto , Eamonn A. Gaffney

Retarded or frequency-dependent hydrodynamic interactions are relevant for velocity relaxation of colloidal particles immersed in a fluid, sufficiently close that their flow patterns interfere. The interactions are also important for…

Fluid Dynamics · Physics 2019-06-26 B. U. Felderhof

Microorganisms are rarely found in Nature swimming freely in an unbounded fluid. Instead, they typically encounter other organisms, hard walls, or deformable boundaries such as free interfaces or membranes. Hydrodynamic interactions between…

Fluid Dynamics · Physics 2013-10-21 Marcelo A. Dias , Thomas R. Powers

We discuss a locomotion of a three-sphere microswimmer in a viscoelastic medium and propose a new type of active microrheology. We derive a relation which connects average swimming velocity and frequency-dependent viscosity of the…

Soft Condensed Matter · Physics 2017-03-30 Kento Yasuda , Ryuichi Okamoto , Shigeyuki Komura

The paper presents a two-phase hydrodynamic model for the numerical simulation of collective motion in a thin layer of active colloids containing spherical microswimmers. The model accounts for three fundamental mechanisms governing the…

Soft Condensed Matter · Physics 2025-12-10 A. Kiverin , S. Luguev , I. Yakovenko

Several recent experiments suggest that rather generally the diffusion of enzymes may be augmented through their activity. We demonstrate that such swimming motility can emerge from the interplay between the enzyme energy landscape and the…

Soft Condensed Matter · Physics 2015-10-08 Xiaoyu Bai , Peter G. Wolynes

The collective swimming of soft robots in an infinite viscous fluid is an emergent phenomenon due to the non-reciprocal hydrodynamic interactions between individual swimmers. These physical interactions give rise to unique spatiotemporal…

Fluid Dynamics · Physics 2024-10-01 R. Pramanik , R. W. C. P. Verstappen , P. R. Onck

The hydrodynamic interactions of a suspension of self-propelled particles are studied using a direct numerical simulation method which simultaneously solves for the host fluid and the swimming particles. A modified version of the "Smoothed…

Soft Condensed Matter · Physics 2013-01-15 John J. Molina , Yasuya Nakayama , Ryoichi Yamamoto