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Bacteria can form a great variety of spatially heterogeneous cell density patterns, ranging from simple concentric rings to dynamical spiral waves appearing in growing colonies. These pattern formation phenomena are important as they…

Populations and Evolution · Quantitative Biology 2025-01-17 John S. Chuang , Riccardo Rao , Stanislas Leibler

Microorganisms living in microfluidic environments often form multi-species swarms, where they can leverage collective motions to achieve enhanced transport and spreading. Nevertheless, there is a general lack of physical understandings of…

Fluid Dynamics · Physics 2022-02-14 Bryce Palmer , Wen Yan , Tong Gao

Pattern formation in active biological matter typically arises from the feedback between chemical concentration fields and mechanical stresses. The actomyosin cortex of cells is an archetypal example of an active thin film that displays…

Biological Physics · Physics 2025-10-17 Jason R. Picardo , V. Jemseena , K. Vijay Kumar

A mixture of spinning particles of two different types represents a system where both nonreciprocity and chirality determine the emergent dynamics. In this work we present a minimal model for a two-species mixture of chiral active…

Statistical Mechanics · Physics 2025-10-27 Giulia Pisegna , Suropriya Saha

One of the most interesting phenomena in the soft-matter realm consists in the spontaneous formation of super-molecular structures (microphases) in condition of thermodynamic equilibrium. A simple mechanism responsible for this…

Soft Condensed Matter · Physics 2017-08-23 A. Imperio , D. Pini , L. Reatto

Living creatures exhibit a remarkable diversity of locomotion mechanisms, evolving structures specialised for interacting with their environment. In the vast majority of cases, locomotor behaviours such as flying, crawling, and running, are…

Biological Physics · Physics 2020-07-28 Kirsty Y. Wan

Cooperation is vital for the survival of a swarm$^1$. Large scale cooperation allows murmuring starlings to outmaneuver preying falcons$^2$, shoaling sardines to outsmart sea lions$^3$, and homo sapiens to outlive their Pleistocene…

Soft Condensed Matter · Physics 2021-01-01 Matan Yah Ben Zion , Alvin Modin , Yaelin Caba , Paul M Chaikin

This article focuses on phase and aggregation behavior for linear chains composed of blocks of hydrophilic and hydrophobic segments. Phase and conformational transitions of patterned chains are relevant for understanding liquid-liquid…

Soft Condensed Matter · Physics 2024-05-15 Athanassios Z. Panagiotopoulos

Active matter comprises individually driven units that convert locally stored energy into mechanical motion. Interactions between driven units lead to a variety of non-equilibrium collective phenomena in active matter. One of such phenomena…

Soft Condensed Matter · Physics 2021-06-15 Mingji Huang , Wensi Hu , Siyuan Yang , Quan-Xing Liu , H. P. Zhang

In their search for metabolic resources microbes swim through viscous environments that present physical anisotropies, including steric obstacles across a wide range of sizes. Hydrodynamic forces are known to significantly alter swimmer…

Soft Condensed Matter · Physics 2020-07-03 Kentaro Hoeger , Tristan Ursell

We propose a simple model of self-propelled particles to show that coherent structures, such as jets and swirls, can arise from a plausible microscopic mechanisms: (i) the elongated shape of the self-propelled particles with (ii) the…

Soft Condensed Matter · Physics 2009-11-13 Nicholas Sambelashvili , A. W. C. Lau , David Cai

In living cells, molecular motors create activity that enhances the diffusion of particles throughout the cytoplasm, and not just ones attached to the motors. We demonstrate initial steps toward creating artificial cells that mimic this…

Soft Condensed Matter · Physics 2019-05-01 Viva R. Horowitz , Zachary C. Chambers , İrep Gözen , Thomas G. Dimiduk , Vinothan N. Manoharan

Despite their vast morphological diversity, many invertebrates have similar larval forms characterized by ciliary bands, innervated arrays of beating cilia that facilitate swimming and feeding. Hydrodynamics suggests that these bands should…

Fluid Dynamics · Physics 2018-04-25 William Gilpin , Vivek N. Prakash , Manu Prakash

We show that activity and broken fore-aft shape symmetry enable microswimmers to cross streamlines in nonuniform shear, a key yet overlooked factor in active cross-stream migration. Using a model of flagellated microswimmers in microchannel…

Soft Condensed Matter · Physics 2025-04-24 Derek C. Gomes , Tapan C. Adhyapak

Disordered hyperuniform structures are an exotic state of matter having vanishing long-wavelength density fluctuations similar to perfect crystals but without long-range order. Although its importance in materials science has been brought…

Soft Condensed Matter · Physics 2019-01-29 Qun-li Lei , Massimo Pica Ciamarra , Ran Ni

We study the structure and phase behavior of a binary mixture where one of the components is self-propelling in nature. The inter-particle interactions in the system were taken from the Asakura-Oosawa model, for colloid-polymer mixtures,…

Statistical Mechanics · Physics 2015-06-17 Subir K. Das , Sergei Egorov , Benjamin Trefz , Peter Virnau , Kurt Binder

We show that a two-dimensional system of flocking microswimmers interacting hydrodynamically can be expressed using a Hamiltonian formalism. The Hamiltonian depends strictly on the angles between the particles and their swimming…

Soft Condensed Matter · Physics 2023-05-23 Yuval Shoham , Naomi Oppenheimer

We experimentally investigate the dynamics of synthetic active particles composed of gravitationally bouncing, superwalking droplets confined within an annular fluid bath. Driven by a topologically pumping dual-frequency waveform, the…

Fluid Dynamics · Physics 2025-12-19 Rudra Sekhri , Rahil N. Valani , Tapio Simula

Active matter has been the object of huge amount of research in recent years for its important fundamental and applicative properties. In this paper we investigate active suspensions of micro-swimmers through direct numerical simulation, so…

Soft Condensed Matter · Physics 2021-02-03 Sergio Chibbaro , Astrid Decoene , Sebastien Martin , Fabien Vergnet

Inspired by the classical Kepler and Rutherford problem, we investigate an analogous set-up in the context of active microswimmers: the behavior of a deformable microswimmer in a swirl flow. First we identify new steady bound states in the…

Soft Condensed Matter · Physics 2014-09-11 Mitsusuke Tarama , Andreas M. Menzel , Hartmut Löwen