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Suspensions of swimming particles exhibit complex collective behaviors driven by hydrodynamic interactions, showing persistent large-scale flows and long-range correlations. While heavily studied, it remains unclear how such structures…

Soft Condensed Matter · Physics 2025-05-27 Bryce Palmer , Scott Weady , Michael O'Brien , Blakesley Burkhart , Michael J. Shelley

Previous particle-based computer simulations have revealed a significantly more pronounced tendency of spontaneous global polar ordering in puller (contractile) microswimmer suspensions than in pusher (extensile) suspensions. We here…

Soft Condensed Matter · Physics 2018-10-23 Christian Hoell , Hartmut Löwen , Andreas M. Menzel

We show that a two-dimensional system of flocking microswimmers interacting hydrodynamically can be expressed using a Hamiltonian formalism. The Hamiltonian depends strictly on the angles between the particles and their swimming…

Soft Condensed Matter · Physics 2023-05-23 Yuval Shoham , Naomi Oppenheimer

In their search for metabolic resources microbes swim through viscous environments that present physical anisotropies, including steric obstacles across a wide range of sizes. Hydrodynamic forces are known to significantly alter swimmer…

Soft Condensed Matter · Physics 2020-07-03 Kentaro Hoeger , Tristan Ursell

The majority of studies on self-propelled particles and microswimmers concentrates on objects that do not feature a deterministic bending of their trajectory. However, perfect axial symmetry is hardly found in reality, and shape-asymmetric…

Statistical Mechanics · Physics 2018-07-18 Christian Hoell , Hartmut Löwen , Andreas M. Menzel

We use numerical simulations to probe the dynamics of concentrated suspensions of spherical microswimmers interacting hydrodynamically. Previous work in the dilute limit predicted orientational instabilities of aligned suspensions for both…

Soft Condensed Matter · Physics 2013-02-12 Arthur A. Evans , Takuji Ishikawa , Takami Yamaguchi , Eric Lauga

We study different types of microswimmers moving in channels with varying cross section and thereby interacting hydrodynamically with the channel walls. Starting from the Smoluchowski equation for a dilute suspension, for which interactions…

Soft Condensed Matter · Physics 2017-05-24 Paolo Malgaretti , Holger Stark

A striking feature of the collective behavior of spherical microswimmers is that for sufficiently strong self-propulsion they phase-separate into a dense cluster coexisting with a low-density dis- ordered surrounding. Extending our previous…

Soft Condensed Matter · Physics 2016-10-31 Johannes Blaschke , Maurice Maurer , Karthik Menon , Andreas Zöttl , Holger Stark

Transport of material across liquid interfaces is ubiquitous for living cells and is also a crucial step in drug delivery and in many industrial processes. The fluids that are present on either side of the interfaces will usually have…

Soft Condensed Matter · Physics 2023-05-24 Chao Feng , John J. Molina , Matthew S. Turner , Ryoichi Yamamoto

We explore the impact of hydrodynamic interactions on the conformational and dynamical properties of wet tangentially-driven active polymers using multiparticle collision dynamics simulations. By analyzing active filaments with varying…

Soft Condensed Matter · Physics 2024-10-24 Loek van Steijn , Mohammad Fazelzadeh , Sara Jabbari-Farouji

Microorganisms can preferentially orient and move along gradients of a chemo-attractant (i.e., chemotax) while colonies of many microorganisms can collectively undergo complex dynamics in response to chemo-attractants that they themselves…

Biological Physics · Physics 2013-10-30 Enkeleida Lushi , Raymond E. Goldstein , Michael J. Shelley

Cooperative motion in biological microswimmers is crucial for their survival as it facilitates adhesion to surfaces, formation of hierarchical colonies, efficient motion, and enhanced access to nutrients. Synthetic microswimmers currently…

Soft Condensed Matter · Physics 2022-04-12 Stefania Ketzetzi , Melissa Rinaldin , Pim Dröge , Joost de Graaf , Daniela J. Kraft

Catalytic microswimmers convert the chemical energy of a fuel into motion, sustaining spatial chemical gradients and fluid flows that drive their propulsion. This leads to unconventional individual behavior and the emergence of collective…

Soft Condensed Matter · Physics 2024-09-04 Celso Carrasco , Quentin Martinet , Zaiyi Shen , Juho S. Lintuvuori , Jérémie Palacci , Antoine Aubret

Particle-particle interactions are of paramount importance in every multi-body system as they determine the collective behaviour and coupling strength. Many well-known interactions like electro-static, van der Waals or screened Coulomb,…

Biological Physics · Physics 2015-07-17 Karolis Misiunas , Stefano Pagliara , Eric Lauga , John R. Lister , Ulrich F. Keyser

The paper presents a two-phase hydrodynamic model for the numerical simulation of collective motion in a thin layer of active colloids containing spherical microswimmers. The model accounts for three fundamental mechanisms governing the…

Soft Condensed Matter · Physics 2025-12-10 A. Kiverin , S. Luguev , I. Yakovenko

We investigate theoretically the collective dynamics of a suspension of low Reynolds number swimmers that are confined to two dimensions by a thin fluid film. Our model swimmer is characterized by internal degrees of freedom which locally…

Biological Physics · Physics 2015-05-19 M. Leoni , T. B. Liverpool

Populations of swimming microorganisms produce fluid motions that lead to dramatically enhanced diffusion of tracer particles. Using simulations of suspensions of swimming particles in a periodic domain, we capture this effect and show that…

Soft Condensed Matter · Physics 2008-07-22 Patrick T. Underhill , Juan P. Hernández-Ortiz , Michael D. Graham

Geometric confinements are frequently encountered in soft matter systems and in particular significantly alter the dynamics of swimming microorganisms in viscous media. Surface-related effects on the motility of microswimmers can lead to…

The general problem of tracer diffusion in non-equilibrium baths is important in a wide range of systems, from the cellular level to geographical lengthscales. In this paper, we revisit the archetypical example of such a system: a…

Soft Condensed Matter · Physics 2023-11-08 Henrik Nordanger , Alexander Morozov , Joakim Stenhammar

Suspensions of rear- and front-actuated microswimmers immersed in a fluid, known respectively as ``pushers'' and ``pullers'', display qualitatively different collective behaviours: beyond a characteristic density, pusher suspensions exhibit…

Soft Condensed Matter · Physics 2020-07-08 Dóra Bárdfalvy , Shan Anjum , Cesare Nardini , Alexander Morozov , Joakim Stenhammar