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Microorganisms are rarely found in Nature swimming freely in an unbounded fluid. Instead, they typically encounter other organisms, hard walls, or deformable boundaries such as free interfaces or membranes. Hydrodynamic interactions between…

Fluid Dynamics · Physics 2013-10-21 Marcelo A. Dias , Thomas R. Powers

We computationally study the kinematics of a simple model reciprocal swimmer (asymmetric dumbbell) as a function of the Reynolds number (Re) and investigate how the onset and gradual increase of inertia impacts the swimming behavior: a…

Soft Condensed Matter · Physics 2020-07-01 Thomas Dombrowski , Daphne Klotsa

During contraction the energy of muscle tissue increases due to energy from the hydrolysis of ATP. This energy is distributed across the tissue as strain-energy potentials in the contractile elements, strain-energy potential from the 3D…

Biological Physics · Physics 2021-01-13 J. M. Wakeling , S. A. Ross , D. S. Ryan , B. Bolsterlee , R. Konno , S. Domínguez , N. Nigam

We use the boundary element method to study the low-Reynolds number locomotion of a spherical model microorganism in a circular tube. The swimmer propels itself by tangen- tial or normal surface motion in a tube whose radius is on the order…

Fluid Dynamics · Physics 2013-06-11 Lailai Zhu , Eric Lauga , Luca Brandt

There is increasing evidence that fish gain energetic benefits when they swim in a school. The most recent indications of such benefits are a lower tail (or fin) beat at the back of a school and reduced oxygen consumption in schooling fish…

Quantitative Methods · Quantitative Biology 2013-07-30 C. K. Hemelrijk , D. A. P. Reid , H. Hildenbrandt , J. T. Padding

Biological locomotion, observed in the flexible wings of birds and insects, bodies and fins of aquatic mammals and fishes, consists of their ability to morph the wings/fins. The morphing capability holds significance in the abilities of…

Fluid Dynamics · Physics 2022-10-19 Pragalbh Dev Singh , Ishan Neogi , Vardhan Niral Shah , Vaibhav Joshi

It has been a long-standing problem how schooling fish optimize their motion by exploiting the vortices shed by the others. A recent experimental study showed that a pair of fish reduce energy consumption by matching the phases of their…

Biological Physics · Physics 2023-12-05 Susumu Ito , Nariya Uchida

We study experimentally a a three-dimensional reduced model of a sail shape performing pitching oscillations around a mean incidence angle ($\alpha_{m}$) with respect to an incoming flow in a hydrodynamic channel at a constant velocity…

Fluid Dynamics · Physics 2026-03-24 Gauthier Bertrand , Ramiro Godoy-Diana , Benjamin Thiria , Marc Fermigier

The dynamics of a triangular magnetocapillary swimmer is studied using the lattice Boltzmann method. Performing extensive numerical simulations taking into account the coupled dynamics of the fluid-fluid interface and of magnetic particles…

We theoretically describe the dynamics of swimmer populations confined in thin liquid films. We first demonstrate that hydrodynamic interactions between confined swimmers only depend on their shape and are independent of their specific…

Soft Condensed Matter · Physics 2015-06-11 Tommaso Brotto , Jean-Baptiste Caussin , Eric Lauga , Denis Bartolo

Tidal and riverine hydrokinetic turbines offer promising solutions for renewable energy generation in aquatic environments. However, their ecological impact, especially on fish behavior, warrants a thorough investigation. This study…

We use particle simulations to reveal two distinct propulsion mechanisms for a scallop-like swimmer to locomote itself in granular media by reciprocally flapping its wings. Based on the discrete element method, we examine the kinematics and…

Soft Condensed Matter · Physics 2025-10-28 Amir Nazemi , Hongyi Xiao

Undulatory locomotion is ubiquitous in nature and observed in different media, from the swimming of flagellated microorganisms in biological fluids, to the slithering of snakes on land, or the locomotion of sandfish lizards in sand. Despite…

Fluid Dynamics · Physics 2017-01-03 Zhiwei Peng , On Shun Pak , Gwynn J. Elfring

We discuss the tidal force, whose notion is sometimes misunderstood in the public domain literature. We discuss the tidal force exerted by a secondary point mass on an extended primary body such as the Earth. The tidal force arises because…

Popular Physics · Physics 2015-06-15 Takuya Matsuda , Hiromu Isaka , Henri M. J. Boffin

When immersed into a fluid of active Brownian particles, passive bodies might start to undergo linear or angular directed motion depending on their shape. Here we exploit the divergence theorem to relate the forces responsible for this…

Soft Condensed Matter · Physics 2021-04-14 Thomas Speck , Ashreya Jayaram

Many biological fluids are composed of suspended polymers immersed in a viscous fluid. A prime example is mucus, where the polymers are also known to form a network. While the presence of this microstructure is linked with an overall…

Fluid Dynamics · Physics 2024-10-10 Adam K. Townsend , Eric E. Keaveny

Biological and artificial microswimmers often have to propel through a variety of environments, ranging from heterogeneous suspending media to strong geometrical confinement. Under confinement, local flow fields generated by microswimmers,…

Soft Condensed Matter · Physics 2024-06-04 Florian A. Overberg , Gerhard Gompper , Dmitry A. Fedosov

Marine organisms manipulate their surrounding flow through their swimming dynamics, which affects the transport of their own odor cues. We demonstrate by direct numerical simulations how a group of swimmers, moving at intermediate Reynolds…

Fluid Dynamics · Physics 2026-05-04 Martin James , Francesco Viola , Agnese Seminara

We study the motion of a microscopic swimmer composed of a semiflexible polymer anchored at the surface of a magnetic sphere using hydrodynamic simulations and scaling arguments. The swimmer is driven by a rotating magnetic field, and…

Soft Condensed Matter · Physics 2007-05-23 A. Alexander-Katz

We present a general theory for determining the force (and torque) exerted on a boundary (or body) in active matter. The theory extends the description of passive Brownian colloids to self-propelled active particles and applies for all…

Soft Condensed Matter · Physics 2023-07-19 Wen Yan , John F. Brady
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