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Animal morphogenesis often involves significant shape changes of epithelial tissue sheets. Great progress has been made in understanding the underlying cellular driving forces and their coordination through biomechanical feedback loops.…

Soft Condensed Matter · Physics 2024-09-23 Nikolas H. Claussen , Fridtjof Brauns

Although cell monolayers typically remain confluent, they can spontaneously develop persistent holes as a result of collective cellular motion. Recent studies on MDCK monolayers cultured on soft substrates have revealed that cells can align…

Soft Condensed Matter · Physics 2025-08-11 Diogo E. P. Pinto , Jan Rozman , Julia M. Yeomans

Eukaryotic cells intrinsically change their shape, by changing the composition of their membrane and by restructuring their underlying cytoskeleton. We present here further studies and extensions of a minimal physical model, describing a…

Soft Condensed Matter · Physics 2023-01-31 Yoav Ravid , Samo Penič , Yuko Mimori-Kiyosue , Shiro Suetsugu , Aleš Iglič , Nir S. Gov

It is widely recognized that the shape of epithelial cells is determined by the tension generated by the actomyosin cortex and the adhesion of cells to the substrate and to each other. To account for these biological and structural…

Biological Physics · Physics 2021-08-11 Nicolas Harmand , Sylvie Hénon

Mechanical forces influence the development and behavior of biological tissues. In many situations these forces are exerted or resisted by elastic compliant structures such as the own-tissue cellular matrix or other surrounding tissues.…

The rheology of biological tissue plays an important role in many processes, from organ formation to cancer invasion. Here, we use a multi-phase field model of motile cells to simulate active microrheology within a tissue monolayer. When…

Soft Condensed Matter · Physics 2022-10-12 Austin Hopkins , Michael Chiang , Benjamin Loewe , Davide Marenduzzo , M. Cristina Marchetti

We theoretically explore fluidization of epithelial tissues by active T1 neighbor exchanges. We show that the geometry of cell-cell junctions encodes important information about the local features of the energy landscape, which we support…

Soft Condensed Matter · Physics 2018-08-31 Matej Krajnc , Sabyasachi Dasgupta , Primož Ziherl , Jacques Prost

Many animal cells change their shape depending on the stiffness of the substrate on which they are cultured: they assume small, rounded shapes in soft ECMs, they elongate within stiffer ECMs, and flatten out on hard substrates. Cells tend…

Cell Behavior · Quantitative Biology 2019-06-24 Elisabeth G. Rens , Roeland M. H. Merks

Cross-sections of cell shapes in a tissue monolayer typically resemble a tiling of convex polygons. Yet, examples exist where the polygons are not convex with curved cell-cell interfaces, as seen in the adaxial epidermis. To date,…

Biological Physics · Physics 2024-06-21 Kyungeun Kim , J. M. Schwarz , Martine Ben Amar

A theoretical model for stratified epithelium is presented. The viscoelastic properties of the tissue is assumed to be dependent on the spatial distribution of proliferative and differentiated cells. Based on this assumption, a hydrodynamic…

Biological Physics · Physics 2016-06-08 Wei-Ting Yeh , Hsuan-Yi Chen

During the spreading of epithelial tissues, the advancing tissue front often develops fingerlike protrusions. Their resemblance to traditional viscous fingering patterns in driven fluids suggests that epithelial fingers could arise from an…

Biological Physics · Physics 2019-03-05 Ricard Alert , Carles Blanch-Mercader , Jaume Casademunt

Constancy of life vitally depends on the internalization of particles through biomembranes. Of particular interest, cellular uptake, including phagocytosis, receptor-mediated endocytosis, and membrane fusion, critically depends on the…

Soft Condensed Matter · Physics 2026-01-07 Amir Khosravanizadeh , Pierre Sens , Farshid Mohammad-Rafiee

Multicellular tissues, such as the epithelium coating a developing embryo, often combine complex tissue shapes with heterogeneity in the spatial arrangement of individual cells. Discrete approximations, such as the cell vertex model, can…

Quantitative Methods · Quantitative Biology 2026-02-25 Oliver E. Jensen , Christopher K. Revell

We analyse a multi-phase field model for an epithelial monolayer with pairwise adhesions between neighbouring cells following an Ornstein-Uhlenbeck process, representing the stochastic turnover of junctional molecular motors. These…

Soft Condensed Matter · Physics 2025-08-27 James N. Graham , Jan Rozman

The coordinated behaviors of epithelial cells are widely observed in tissue development, such as re-epithelialization, tumor growth, and morphogenesis. In these processes, cells either migrate collectively or organize themselves into…

Soft Condensed Matter · Physics 2022-08-30 Tiankai Zhao , Hongyan Yuan

Cell neighbor exchanges play a critical role in regulating tissue fluidity during epithelial morphogenesis and repair. In vivo, these neighbor exchanges are often hindered by the formation of transiently stable four-fold vertices, which can…

Biological Physics · Physics 2023-10-17 Fernanda Pérez-Verdugo , Shiladitya Banerjee

Cells in an epithelial monolayer coordinate motion with their neighbors giving rise to collectively moving packs of sizes spanning multiple cell diameters. The physical mechanism controlling the pack size, however, remains unclear. A…

Biological Physics · Physics 2021-07-21 Aashrith Saraswathibhatla , Silke Henkes , Emmett E. Galles , Rastko Sknepnek , Jacob Notbohm

Living cells are soft bodies of a characteristic form, but endowed with a capacity for a steady turnover of their structures. Both of these material properties, i.e. recovery of the shape after an external stress has been imposed and…

Soft Condensed Matter · Physics 2007-05-23 Erwin Frey , Klaus Kroy , Jan Wilhelm

The alignment of fibers and cells in living tissues affect their mechanical properties and functionality. In this context, one can draw an analogy between tissues and nematic liquid crystal elastomers. We explore this analogy by growing…

Soft Condensed Matter · Physics 2025-10-08 Kirsten Endresen , Aniruddh Murali , Birte Geerds , Daniel J. G. Pearce , Francesca Serra

The conditions under which biological cells switch from a static to a motile state are fundamental to the understanding of many healthy and pathological processes. In this paper, we show that even in the presence of a fully symmetric…

Biological Physics · Physics 2024-01-12 Jocelyn Étienne , Pierre Recho
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