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Adhesive cell-substrate interactions are crucial for cell motility and are responsible for the necessary traction that propels cells. These interactions can also change the shape of the cell, analogous to liquid droplet wetting on adhesive…

Experiments and theory have shown that cell monolayers and epithelial tissues exhibit solid-liquid and glass-liquid transitions. These transitions are biologically relevant to our understanding of embryonic development, wound healing, and…

Soft Condensed Matter · Physics 2020-07-22 Benjamin Loewe , Michael Chiang , Davide Marenduzzo , M. Cristina Marchetti

Coordinated and cooperative motion of cells is essential for embryonic development, tissue morphogenesis, wound healing and cancer invasion. A predictive understanding of the emergent mechanical behaviors in collective cell motion is…

Biological Physics · Physics 2022-10-04 Michael F. Staddon , Michael P. Murrell , Shiladitya Banerjee

Growing experimental evidence indicates that topological defects could serve as organizing centers in the morphogenesis of tissues. Here, we provide a quantitative explanation for this phenomenon, rooted in the buckling theory of deformable…

Soft Condensed Matter · Physics 2022-04-21 Ludwig A. Hoffmann , Livio Nicola Carenza , Julia Eckert , Luca Giomi

Living tissues undergo wetting transitions: On a surface, they can either form a droplet-like cell aggregate or spread as a monolayer of migrating cells. Tissue wetting depends not only on the chemical but also on the mechanical properties…

Biological Physics · Physics 2020-05-14 Ricard Alert , Jaume Casademunt

The complex spatiotemporal flow patterns in living tissues, driven by active forces, have many of the characteristics associated with inertial turbulence even though the Reynolds number is extremely low. Analyses of experimental data from…

Soft Condensed Matter · Physics 2022-11-29 Xin Li , Sumit Sinha , T. R. Kirkpatrick , D. Thirumalai

Rigidity is an emergent property of materials - it is not a feature of individual components that comprise the structure, but instead arises from interactions between many constituent parts. Recently, it has been recognized that…

Soft Condensed Matter · Physics 2025-08-27 Kelly Aspinwall , Tyler Hain , M. Lisa Manning

The rheology of biological tissue is key to processes such as embryo development, wound healing and cancer metastasis. Vertex models of confluent tissue monolayers have uncovered a spontaneous liquid-solid transition tuned by cell shape;…

Soft Condensed Matter · Physics 2023-10-02 Suzanne M. Fielding , James O. Cochran , Junxiang Huang , Dapeng Bi , M. Cristina Marchetti

Coordinated cellular movements are key processes in tissue morphogenesis. Using a cell-based modeling approach we study the dynamics of epithelial layers lining surfaces with constant and varying curvature. We demonstrate that extrinsic…

Soft Condensed Matter · Physics 2024-03-21 Lea Happel , Axel Voigt

Liquid-liquid phase separation is an important mechanism for compartmentalizing the cell's cytoplasm, allowing the dynamic organization of the components necessary for survival. However, it is not clear how phase separation is affected by…

Soft Condensed Matter · Physics 2022-01-31 Estefania Vidal-Henriquez , David Zwicker

How can dense biological tissue maintain sharp boundaries between coexisting cell populations? We explore this question within a simple vertex model for cells, focusing on the role of topology and tissue surface tension. We show that the…

Soft Condensed Matter · Physics 2018-02-07 Daniel M. Sussman , J. M. Schwarz , M. Cristina Marchetti , M. Lisa Manning

The rheology of biological tissue plays an important role in many processes, from organ formation to cancer invasion. Here, we use a multi-phase field model of motile cells to simulate active microrheology within a tissue monolayer. When…

Soft Condensed Matter · Physics 2022-10-12 Austin Hopkins , Michael Chiang , Benjamin Loewe , Davide Marenduzzo , M. Cristina Marchetti

During embryonic morphogenesis, tissues undergo dramatic deformations in order to form functional organs. Similarly, in adult animals, living cells and tissues are continually subjected to forces and deformations. Therefore, the success of…

Soft Condensed Matter · Physics 2024-05-06 Michael J. Hertaeg , Suzanne M. Fielding , Dapeng Bi

The mechanical properties of biological materials are spatially heterogeneous. Typical tissues are made up of a spanning fibrous extracellular matrix in which various inclusions, such as living cells, are embedded. While the influence of…

Soft Condensed Matter · Physics 2025-09-04 Jordan L. Shivers , Jingchen Feng , Fred C. MacKintosh

Cellular decision making allows cells to assume functionally different phenotypes in response to microenvironmental cues, without genetic change. It is an open question, how individual cell decisions influence the dynamics at the tissue…

We formulate a hydrodynamic theory of confluent epithelia: i.e. monolayers of epithelial cells adhering to each other without gaps. Taking advantage of recent progresses toward establishing a general hydrodynamic theory of p-atic liquid…

Soft Condensed Matter · Physics 2024-01-11 Josep-Maria Armengol-Collado , Livio Nicola Carenza , Luca Giomi

Complex tissue flows in epithelia are driven by intra- and inter-cellular processes that generate, maintain, and coordinate mechanical forces. There has been growing evidence that cell shape anisotropy, manifested as nematic order, plays an…

Soft Condensed Matter · Physics 2024-12-17 Jan Rozman , Chaithanya K. V. S. , Julia M. Yeomans , Rastko Sknepnek

Vesicle fusion is an important process underlying cell division, transport, and membrane trafficking. In phospholipid systems, a range of fusogens including divalent cations and depletants have been shown to induce adhesion, hemifusion, and…

Soft Condensed Matter · Physics 2023-05-03 Ziyan Fan , Yaam Deckel , Lauren A. Lowe , Daniel W. K. Loo , Tetsuya Yomo , Jack W. Szostak , Collin Nisler , Anna Wang

Connecting cell behavior to tissue shape and mechanics is a key challenge in the physics of morphogenesis. Cytoskeletal turnover precludes a fixed reference state, and tensions are actively generated independently of strain; so conventional…

Soft Condensed Matter · Physics 2026-05-28 Nikolas H. Claussen , Fridtjof Brauns , Boris I. Shraiman

We study the influence of cell-level mechanical heterogeneity in epithelial tissues using a vertex-based model. Heterogeneity in single cell stiffness is introduced as a quenched random variable in the preferred shape index($p_0$) for each…

Biological Physics · Physics 2019-08-07 Xinzhi Li , Amit Das , Dapeng Bi