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The Colless index for bifurcating phylogenetic trees, introduced by Colless (1982), is defined as the sum, over all internal nodes $v$ of the tree, of the absolute value of the difference of the sizes of the clades defined by the children…

Populations and Evolution · Quantitative Biology 2020-07-30 Tomás M. Coronado , Arnau Mir , Francesc Rosselló

Measures of tree balance play an important role in the analysis of phylogenetic trees. One of the oldest and most popular indices in this regard is the Colless index for rooted bifurcating trees, introduced by Colless (1982). While many of…

Populations and Evolution · Quantitative Biology 2020-02-18 Tomás M. Coronado , Mareike Fischer , Lina Herbst , Francesc Rosselló , Kristina Wicke

Measures of tree balance play an important role in various research areas, for example in phylogenetics. There they are for instance used to test whether an observed phylogenetic tree differs significantly from a tree generated by the Yule…

Combinatorics · Mathematics 2019-07-23 Mareike Fischer , Lina Herbst , Kristina Wicke

The Sackin and Colless indices are two widely-used metrics for measuring the balance of trees and for testing evolutionary models in phylogenetics. This short paper contributes two results about the Sackin and Colless indices of trees. One…

Populations and Evolution · Quantitative Biology 2022-07-20 Gary Goh , Michael Fuchs , Louxin Zhang

We define a new balance index for rooted phylogenetic trees based on the symmetry of the evolutive history of every set of 4 leaves. This index makes sense for multifurcating trees and it can be computed in time linear in the number of…

Populations and Evolution · Quantitative Biology 2019-03-25 Tomás M. Coronado , Arnau Mir , Francesc Rosselló , Gabriel Valiente

For two decades, the Colless index has been the most frequently used statistic for assessing the balance of phylogenetic trees. In this article, this statistic is studied under the Yule and uniform model of phylogenetic trees. The main tool…

Probability · Mathematics 2007-05-23 Michael G. B. Blum , Olivier François , Svante Janson

(Im)balance indices can be used to quantify the (im)balance of trees by assigning numerical scores to them. An easy way to generate a new index is to construct a compound index, e.g., a linear combination of established indices. Two of the…

Populations and Evolution · Quantitative Biology 2026-04-20 Linda Knüver , Mareike Fischer

Tree shape statistics, particularly measures of tree (im)balance, play an important role in the analysis of the shape of phylogenetic trees. With applications ranging from testing evolutionary models to studying the impact of fertility…

Populations and Evolution · Quantitative Biology 2024-10-15 Sophie J. Kersting , Kristina Wicke , Mareike Fischer

Several indices that measure the degree of balance of a rooted phylogenetic tree have been proposed so far in the literature. In this work we define and study a new index of this kind, which we call the total cophenetic index: the sum, over…

Populations and Evolution · Quantitative Biology 2014-02-11 Arnau Mir , Francesc Rossello , Lucia Rotger

Tree balance plays an important role in various research areas in phylogenetics and computer science. Typically, it is measured with the help of a balance index or imbalance index. There are more than 25 such indices available, recently…

Populations and Evolution · Quantitative Biology 2025-08-20 Sean Cleary , Mareike Fischer , Katherine St. John

The Colless index is one of the oldest and most widely used balance indices for rooted bifurcating trees. Despite its popularity, its minimum value on the space $\mathcal{T}_n$ of rooted bifurcating trees with $n$ leaves is only known when…

Populations and Evolution · Quantitative Biology 2019-07-24 Tomás M. Coronado , Francesc Rosselló

Full binary trees naturally represent commutative non-associative products. There are many important examples of these products: finite-precision floating-point addition and NAND gates, among others. Balance in such a tree is highly…

Discrete Mathematics · Computer Science 2021-08-27 Laura Monroe

Measures of tree balance play an important role in many different research areas such as mathematical phylogenetics or theoretical computer science. Typically, tree balance is quantified by a single number which is assigned to the tree by a…

Populations and Evolution · Quantitative Biology 2025-07-31 Mareike Fischer , Tom Niklas Hamann , Kristina Wicke

Binary rooted trees, both in the ordered and in the un-ordered case, are well studied structures in the field of combinatorics. The aim of this work is to study particular patterns in these classes of trees. We consider completely…

Combinatorics · Mathematics 2013-03-12 Filippo Disanto

Measures of phylogenetic balance, such as the Colless and Sackin indices, play an important role in phylogenetics. Unfortunately, these indices are specifically designed for phylogenetic trees, and do not extend naturally to phylogenetic…

Populations and Evolution · Quantitative Biology 2022-11-14 François Bienvenu , Gabriel Cardona , Celine Scornavacca

We compute an explicit formula for the expected value of the Colless index of a phylogenetic tree generated under the Yule model, and an explicit formula for the expected value of the Sackin index of a phylogenetic tree generated under the…

Populations and Evolution · Quantitative Biology 2012-01-19 Arnau Mir , Francesc Rossello

Phylogenetic networks play an important role in evolutionary biology as, other than phylogenetic trees, they can be used to accommodate reticulate evolutionary events such as horizontal gene transfer and hybridization. Recent research has…

Populations and Evolution · Quantitative Biology 2024-03-19 Linda Knüver , Mareike Fischer , Marc Hellmuth , Kristina Wicke

The Sackin index is an important measure for the balance of phylogenetic trees. We investigate two extensions of the Sackin index to the class of galled trees and two of its subclasses (simplex galled trees and normal galled trees) where we…

Populations and Evolution · Quantitative Biology 2025-02-21 Michael Fuchs , Bernhard Gittenberger

Tree balance plays an important role in phylogenetics and other research areas, which is why several indices to measure tree balance have been introduced over the years. Nevertheless, a formal definition of what a balance index actually is…

Populations and Evolution · Quantitative Biology 2023-11-10 Mareike Fischer , Lina Herbst , Sophie Kersting , Luise Kühn , Kristina Wicke

Phylogenetic trees are important tools in the study of evolutionary relationships between species. Measures such as the index of Sackin, Colless, and Total Cophenetic have been extensively used to quantify tree balance, one key property of…

Populations and Evolution · Quantitative Biology 2020-09-01 T. Araújo Lima , Marcus A. M. de Aguiar
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