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Recent experiments on the green alga Chlamydomonas that swims using synchronized beating of a pair of flagella have revealed that it exhibits a run-and-tumble behavior similar to that of bacteria such as E. Coli. Using a simple purely…

Cell Behavior · Quantitative Biology 2015-06-12 Rachel R. Bennett , Ramin Golestanian

Synchronization induced by long-range hydrodynamic interactions is attracting attention as a candidate mechanism behind coordinated beating of cilia and flagella. Here we consider a minimal model of hydrodynamic synchronization in the low…

Soft Condensed Matter · Physics 2012-11-26 Nariya Uchida , Ramin Golestanian

In a recent letter (Friedrich et al., Phys. Rev. Lett. 109:138102, 2012), a minimal model swimmer was proposed that propels itself at low Reynolds numbers by a revolving motion of a pair of spheres. The motion of the two spheres can…

Cell Behavior · Quantitative Biology 2015-06-12 Katja Polotzek , Benjamin M. Friedrich

Some types of bacteria use rotating helical flagella to swim. The motion of such organisms takes place in the regime of low Reynolds numbers where viscous effects dominate and where the dynamics is governed by hydrodynamic interactions.…

Soft Condensed Matter · Physics 2007-05-23 M. Reichert , H. Stark

Cilia and flagella exhibit regular bending waves that perform mechanical work on the surrounding fluid, to propel cellular swimmers and pump fluids inside organisms. Here, we quantify a force-velocity relationship of the beating flagellum,…

Cell Behavior · Quantitative Biology 2016-12-21 Gary S. Klindt , Christian Ruloff , Christian Wanger , Benjamin M. Friedrich

Cellular appendages conferring motility, such as flagella or cilia, are known to synchronise their periodic beats. The origin of synchronisation is a combination of long-range hydrodynamic interactions with physical mechanisms allowing the…

Biological Physics · Physics 2021-04-21 Ivan Tanasijević , Eric Lauga

Some microorganisms, such as spermatozoa, synchronize their flagella when swimming in close proximity. Using a simplified model (two infinite, parallel, two-dimensional waving sheets), we show that phase-locking arises from hydrodynamics…

Biological Physics · Physics 2009-09-15 Gwynn J. Elfring , Eric Lauga

We survey the theory synchronization in collections of noisy oscillators. This framework is applied to flagellar synchronization by hydrodynamic interactions. The time-reversibility of hydrodynamics at low Reynolds numbers prompts swimming…

Cell Behavior · Quantitative Biology 2016-11-23 B. M. Friedrich

Experimental studies have demonstrated that spermatozoa synchronize their flagella when swimming in close proximity. In a Newtonian fluid, it was shown theoretically that such synchronization arises passively due to hydrodynamic forces…

Fluid Dynamics · Physics 2010-05-02 Gwynn J. Elfring , On Shun Pak , Eric Lauga

A system of active colloidal particles driven by harmonic potentials to oscillate about the vertices of a regular polygon, with hydrodynamic coupling between all particles, is described by a piece-wise linear model which exhibits various…

Statistical Mechanics · Physics 2013-05-30 Giovanni M. Cicuta , Enrico Onofri , Marco Cosentino Lagomarsino , Pietro Cicuta

Motivated by the observed coordination of nearby beating cilia, we use a scale model experiment to show that hydrodynamic interactions can cause synchronization between rotating paddles driven at constant torque in a very viscous fluid.…

Soft Condensed Matter · Physics 2010-05-26 Bian Qian , Hongyuan Jiang , David A. Gagnon , Kenneth S. Breuer , Thomas R. Powers

The motility of microorganisms is influenced greatly by their hydrodynamic interactions with the fluidic environment they inhabit. We show by direct experimental observation of the bi-flagellated alga Chlamydomonas reinhardtii that fluid…

Biological Physics · Physics 2015-11-03 Boyang Qin , Arvind Gopinath , Jing Yang , Jerry P Gollub , Paulo E Arratia

When swimming in close proximity, some microorganisms such as spermatozoa synchronize their flagella. Previous work on swimming sheets showed that such synchronization requires a geometrical asymmetry in the flagellar waveforms. Here we…

Fluid Dynamics · Physics 2011-08-31 Gwynn J. Elfring , Eric Lauga

Using multiparticle collision dynamics simulations, we investigate the swimming dynamics, orientational behavior, and hydrodynamic interactions of a model swimmer designed to mimic the isolated flagellar apparatus ($FA$) of Chlamydomonas…

Biological Physics · Physics 2025-07-01 Sai Venkata Ramana Ambadipudi , Albert Bae , Azam Gholami

Many active biological particles, such as swimming microorganisms or motor-proteins, do work on their environment by going though a periodic sequence of shapes. Interactions between particles can lead to the phase-synchronization of their…

Soft Condensed Matter · Physics 2022-07-26 Brato Chakrabarti , Michael J. Shelley , Sebastian Fürthauer

The beating flagella of the green alga Chlamydomonas reinhardtii play a prominent role in cellular mechanics, enabling cells to both displace and sense surrounding fluid. Specifically, flagellum-induced fluid transport enables microalgae to…

Fluid Dynamics · Physics 2026-03-10 Md Iftekhar Yousuf Emon , Gregorius R. Pradipta , Xiang Cheng , Xin Yong

Microorganisms swimming through viscous fluids imprint their propulsion mechanisms in the flow fields they generate. Extreme confinement of these swimmers between rigid boundaries often arises in natural and technological contexts, yet…

Soft Condensed Matter · Physics 2022-01-26 Debasmita Mondal , Ameya G. Prabhune , Sriram Ramaswamy , Prerna Sharma

Hydrodynamic coupling of flexible flags in axial flows may profoundly influence their flapping dynamics, in particular driving their synchronization. This work investigates the effect of such coupling on the harvesting efficiency of coupled…

Fluid Dynamics · Physics 2016-09-13 Yifan Xia , Olivier Doare , Sebastien Michelin

Flagella allow eukaryotic cells to move and pump fluid. We present the first three-dimensional, time-resolved imaging of the flagellar waveform of Chlamydomonas reinhardtii, a model alga found in fresh water. During the power stroke, we…

Biological Physics · Physics 2022-10-18 Laurence G. Wilson , Martin A. Bees

Living creatures exhibit a remarkable diversity of locomotion mechanisms, evolving structures specialised for interacting with their environment. In the vast majority of cases, locomotor behaviours such as flying, crawling, and running, are…

Biological Physics · Physics 2020-07-28 Kirsty Y. Wan