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When swimming at low Reynolds numbers, inertial effects are negligible and reciprocal movements cannot induce net motion. Instead, symmetry breaking is necessary to achieve net propulsion. Directed swimming can be supported by magnetic…

Soft Condensed Matter · Physics 2026-03-31 Theo Lequy , Andreas M. Menzel

A single flexible filament can be actuated to escape from the scallop theorem and generate net propulsion at low Reynolds number. In this work, we study the dynamics of a simple boundary-driven multi-filament swimmer, a two-arm clamshell…

Fluid Dynamics · Physics 2022-03-07 Shiyuan Hu , Jun Zhang , Michael J. Shelley

Shear-thinning is an important rheological property of many biological fluids, such as mucus, whereby the apparent viscosity of the fluid decreases with shear. Certain microscopic swimmers have been shown to progress more rapidly through…

Fluid Dynamics · Physics 2013-09-06 Thomas D. Montenegro-Johnson , Daniel Loghin , David J. Smith

Swimming and pumping at low Reynolds numbers are subject to the "Scallop theorem", which states that there will be no net fluid flow for time reversible motions. Living organisms such as bacteria and cells are subject to this constraint,…

Soft Condensed Matter · Physics 2009-01-20 M. Leoni , J. Kotar , B. Bassetti , P. Cicuta , M. Cosentino Lagomarsino

From bacteria and sperm cells to artificial microrobots, self-propelled microscopic objects at low Reynolds numbers often perceive fluctuating mechanical and chemical stimuli and contact exterior wall boundaries both in nature and the…

Soft Condensed Matter · Physics 2023-12-22 Yoshiki Hiruta , Kenta Ishimoto

It has been argued that an extended, quasi-rigid body evolving freely in curved spacetime can deviate from its natural trajectory by simply performing cyclic deformations. More interestingly, in the limit of rapid cycles, the amount of…

General Relativity and Quantum Cosmology · Physics 2017-03-28 Rodrigo Andrade e Silva , George E. A. Matsas , Daniel A. T. Vanzella

We analyze a minimal model for a rigid spherical microswimmer and explore the consequences of its extended surface on the interplay between its self-propulsion and flow properties. The model is the first order representation of…

Soft Condensed Matter · Physics 2017-12-06 Tapan Chandra Adhyapak , Sara Jabbari-Farouji

We review recent work on active colloids or swimmers, such as self-propelled microorganisms, phoretic colloidal particles, and artificial micro-robotic systems, moving in fluid-like environments. These environments can be water-like and…

Biological Physics · Physics 2016-02-09 Alison E. Patteson , Arvind Gopinath , Paulo E. Arratia

The biological fluids encountered by self-propelled cells display complex microstructures and rheology. We consider here the general problem of low-Reynolds number locomotion in a complex fluid. {Building on classical work on the transport…

Fluid Dynamics · Physics 2014-10-16 Eric Lauga

Self-propulsion at low Reynolds number is notoriously restricted, a concept that is commonly known as the "scallop theorem". Here we present a truly self-propelled swimmer (force- and torque- free) that, while unable to swim in a Newtonian…

Fluid Dynamics · Physics 2021-11-23 Laurel A. Kroo , Jeremy P. Binagia , Noah Eckman , Manu Prakash , Eric S. G. Shaqfeh

By synergistically combining modeling, simulation and experiments, we show that there exists a regime of self-propulsion in which the inertia in the fluid dynamics can be separated from that of the swimmer. This is demonstrated by the…

Many microorganisms swim through gels and non-Newtonian fluids in their natural environments. In this paper, we focus on microorganisms which use flagella for propulsion. We address how swimming velocities are affected in nonlinearly…

Biological Physics · Physics 2010-04-07 Henry C. Fu , Charles W. Wolgemuth , Thomas R. Powers

We experimentally study a scallop-like swimmer with reciprocally flapping wings in a nearly frictionless, cohesive granular medium consisting of hydrogel spheres. Significant locomotion is found when the swimmer's flapping frequency matches…

Soft Condensed Matter · Physics 2026-04-29 Hongyi Xiao , Jing Wang , Achim Sack , Ralf Stannarius , Thorsten Pöschel

We illustrate a concept for shape-changing microswimmers, which exploits the hysteresis of a shape transition of an elastic object, by an elastic disk undergoing cyclic localized swelling. Driving the control parameter of a hysteretic shape…

Soft Condensed Matter · Physics 2020-12-29 Christian Wischnewski , Jan Kierfeld

An articulated body is defined as a finite number of rigid bodies connected by a set of arbitrary constraints that limit the relative motion between pairs of bodies. Such a general definition encompasses a wide variety of situations in the…

Computational Physics · Physics 2024-07-12 Florencio Balboa Usabiaga , Blaise Delmotte

Viscoelastic fluids impact the locomotion of swimming microorganisms and can be harnessed to devise new types of self-propelling devices. Here we report on experiments demonstrating the use of normal stress differences for propulsion. Rigid…

Fluid Dynamics · Physics 2020-12-10 Jhonny A. Puente-Velazquez , Francisco A. Godinez , Eric Lauga , Roberto Zenit

Hydrodynamic interactions are crucial for determining the cooperative behavior of microswimmers at low Reynolds numbers. Here we provide a comprehensive analysis of the scaling and strength of the interactions in the case of a pair of…

Soft Condensed Matter · Physics 2021-02-11 Sebastian Ziegler , Thomas Scheel , Maxime Hubert , Jens Harting , Ana-Sunčana Smith

Predicting how a deformable body moves and deforms in a viscous flow underlies problems ranging from microorganism locomotion to soft microrobotics, yet existing frameworks are either problem-specific or ill-suited to inverse design. We…

Fluid Dynamics · Physics 2026-05-25 Christophe Eloy

Most classical work on the hydrodynamics of low-Reynolds-number swimming addresses deterministic locomotion in quiescent environments. Thermal fluctuations in fluids are known to lead to a Brownian loss of the swimming direction. As most…

Fluid Dynamics · Physics 2014-06-18 Mario Sandoval , Navaneeth K. M. , Ganesh Subramanian , Eric Lauga

We theoretically and computationally study the low-Reynolds-number hydrodynamics of a linear active microswimmer surfing on a compressible thin fluid layer characterized by an odd viscosity. Since the underlying three-dimensional fluid is…

Soft Condensed Matter · Physics 2023-10-25 Yuto Hosaka , Ramin Golestanian , Abdallah Daddi-Moussa-Ider
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